[HN Gopher] Why do bees die when they sting you? (2021)
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       Why do bees die when they sting you? (2021)
        
       Author : ohjeez
       Score  : 331 points
       Date   : 2025-01-18 15:32 UTC (1 days ago)
        
 (HTM) web link (www.subanima.org)
 (TXT) w3m dump (www.subanima.org)
        
       | tibbon wrote:
       | The group selection part is really interesting evolution-wise. It
       | seems a very slow and difficult method of selection. I had never
       | considered how something dying, and not passing along their
       | genetics, could enforce a genetic trait.
        
         | odyssey7 wrote:
         | It requires such a depth of evolution as to make it absurd to
         | imagine that the genus homo is the point at which altruism
         | emerged. Animals care about each other.
        
         | crazydoggers wrote:
         | "Group selection" is not a thing. The article hand waves this
         | always with
         | 
         | > some biologists still get really triggered about group
         | selection and deny its evolutionary importance
         | 
         | Which is dishonest at best. The _vast_ majority of biologist
         | have realized group selection doesn't work as proposed. [0]
         | 
         | What people thought was group selection was just kin selection
         | working over time.
         | 
         | All evolution works at the level of the gene. Genes "want" to
         | reproduce more of themselves. And if the same gene is in a kin,
         | then it can favor enhancing the survival of kin that carry
         | copies of itself. At a macro level this can be misreported as
         | group selection, but to be sure, the selection is happening at
         | the level of the gene, and reaches at most to kin sharing
         | genes.
         | 
         | The article then goes on to say
         | 
         | > The nice story I told above about the evolution of altruism
         | could just have easily been applied to humans. Yet we do not
         | exist in eusocial colonies, so there must be something else
         | going on
         | 
         | And he then talks about gene selection and the fact that bees
         | are haplodiploidy, which is indeed the cause of the "altruism"
         | we see.
         | 
         | His dismissal of haplodiploidy at the end of the article is a
         | weak argument. Just because haplodiploidy in other species
         | doesn't lead to eusocial groups, or that eusocial groups can
         | occur without haplodiploidy are not sufficient arguments that
         | dismiss the effects of haplodiploidy and kin selection favoring
         | altruism in eusocial bees.
         | 
         | I highly recommend people interested in these topics to read
         | the seminal _Selfish Gene_ by Richard Dawkins. [1]
         | 
         | 0. https://en.m.wikipedia.org/wiki/Group_selection#Criticism
         | 
         | 1. https://en.m.wikipedia.org/wiki/The_Selfish_Gene
        
           | glenstein wrote:
           | Agreed, it's a disappointing and discrediting detour in an
           | article that's about a fascinating topic. As you note, this
           | has been worked out via haplodiploidy, which doesn't require
           | venturing into theorizing about group selection or altruism.
           | 
           | And just to take a beat, and explain why group selection
           | "triggers" people (in the author's wording), it's because it
           | violates our fundamental, bedrock idea of causality which is
           | no small thing, and anyone having a cavalier attitude about
           | that probably doesn't belong in a room where these ideas are
           | being deliberated. We understand physics to be causally
           | closed, and expect "higher level" explanations to be
           | compatible with the constraints of physics.
           | 
           | A model example in taking causality seriously, and proceeding
           | with extreme care and extreme caution about challenging that
           | intuition, I think is best exhibited in Quantum Mechanics,
           | where, after excruciatingly careful examination of data and
           | lots of hard thinking about implications, and lots of
           | accounting for it's almost vulgar challenge to our
           | intuitions, do we dare offer a model that challenges our
           | basic idea of causation. That deviation is appropriately
           | treated as profound, by contrast with the fast and loose
           | invocation of group selection you find in some evolutionary
           | explanations.
        
             | crazydoggers wrote:
             | Yes! To put a finer point on it, group selection theories
             | don't have a specific physical explanation for how they
             | operate, instead veering into philosophical explanations.
             | 
             | Ultimately natural selection must operate on the gene.
             | Genes are the only source of information that gets passed
             | to offspring through germ line cells in sexual reproduction
             | or mitosis in asexual reproduction (don't get me started on
             | the fad of epigenetics, which is just a fancy term for
             | standard DNA controlled embryological differentiation.)
             | 
             | The replication of genes and the information they encode,
             | are the physical cause of the effect of phenotypes.
             | 
             | Group selection theorists (of which there are few) have no
             | physical cause that allows selection to occur on the level
             | of the group, and there's no sound hypothesis of such that
             | I have heard of. You'd need some physical mechanism for
             | information flow between individuals in a group for that to
             | be the case, and outside of kin inheritance, there's
             | nothing like that that exists.
        
         | hbn wrote:
         | Think about how most people are naturally scared of heights, or
         | snakes. A lot of dogs get freaked out by snakes too, or if you
         | play with a hair clip in front of them, which looks like a
         | snake.
         | 
         | The ones that aren't afraid of those things are more likely to
         | die from falling off a cliff or being injected with venom.
         | 
         | I'm personally someone who is freaked out heavily by insects. I
         | know logically a house centipede or a harmless spider can't
         | hurt me, but seemingly my brain has something in it that
         | overrides my entire body when I see one and disgusts me.
         | Usually it's irrational, but it probably helps humanity on a
         | larger scale to avoid the ones that are dangerous!
         | 
         | There's a lot about the humans body that naturally gives us
         | non-logical instincts that help us to survive and breed. People
         | like having sex, regardless of whether they want a baby.
         | There's no logic to it, but we know what we like!
         | 
         | The more advanced we get, the more it becomes apparent that
         | we're just monkeys in shoes.
        
           | caseyohara wrote:
           | > A lot of dogs get freaked out by snakes too, or if you play
           | with a hair clip in front of them, which looks like a snake.
           | 
           | Some cats are afraid of cucumbers, presumably because the
           | shape and color resembles a snake. Here's a funny
           | compilation: https://youtu.be/oDpQ2uGLUKU
        
             | FartyMcFarter wrote:
             | > Some cats are afraid of cucumbers, presumably because the
             | shape and color resembles a snake. Here's a funny
             | compilation: https://youtu.be/oDpQ2uGLUKU
             | 
             | It's funny in a way, but if you think about it it's
             | actually abusive.
             | 
             | Would you think it's funny if you were terrified of snakes
             | and someone randomly put a fake snake next to you when you
             | were just relaxing?
        
         | s1artibartfast wrote:
         | group selection works a lot better when the sacrificing
         | individuals are sterile, with no other hope of passing on their
         | genetics.
         | 
         | See also Eunuchs and Castration as a way to recreate a similar
         | dynamic with humans. Castration had the fascinating ability to
         | bind the interests of the Eunuchs more closely with the power
         | structures and rules, by removing the option of family and
         | progeny of their own.
         | 
         | https://en.wikipedia.org/wiki/Eunuch#Asia_and_Africa
         | 
         | [Edit] As crazydoggers points out, it is probably better to
         | view this through the lens of kin selection, with reproducers
         | as the evolutionary agents.
         | 
         | https://news.ycombinator.com/item?id=42749677
        
         | DontchaKnowit wrote:
         | Is that not essentially the only way that selection happens?
         | You are just desvribing basic natural selection
        
         | gus_massa wrote:
         | It's somewhat explained near the end of the article. Sex in
         | bees (and ants) is weird. https://en.wikipedia.org/wiki/Sex-
         | determination_system
         | 
         | Humans use XY system, so we share 50% of your genes with your
         | children, parents and siblings (in average).
         | 
         | Bees and ants use X0 system. A female bee share 50% of your
         | genes with their own daughters, 75% with their mother and 75%
         | with their female parents and 75% with their female siblings
         | (in average).
         | 
         | So, from the bee's genes point of view instead of having their
         | own children it's better to kidnap their mother and force her
         | to have more female children. And a consequence is that instead
         | of running away to form their new family in a safe place it's
         | better to die protecting their mother.
        
           | JadeNB wrote:
           | > 75% with their mother and 75% with their female parents
           | 
           | Are these different things for bees?
        
             | gus_massa wrote:
             | Sorry, cut&paste typo.
        
           | penteract wrote:
           | Your conclusion is right, but in bees, sex is determined by
           | Haplodiploidy ( https://en.wikipedia.org/wiki/Haplodiploidy
           | ), not X0. Also, the daughters have the same number of
           | chromosomes as the mother so they share 50%, not 75% of their
           | genes with their mother (they do share an average of 75% of
           | their genes with their sisters).
        
       | redundantly wrote:
       | The next time my wife asks me why we have something new, my
       | response will be "Because in a capitalistic society, you can
       | exchange money for ____."
        
         | GauntletWizard wrote:
         | Money can be exchanged for goods and services       - Homer
         | Simpson's brain
         | 
         | https://youtu.be/A81DYZh6KaQ
        
         | ahazred8ta wrote:
         | There are many valid explanations.
         | https://www.ft.com/lol-404-theories
        
           | idatum wrote:
           | This link is awesome. Thanks for sharing!
        
       | raldi wrote:
       | > the result is the picture at the top of this article
       | 
       | But there is no picture at the top of the article, at least on
       | mobile.
        
         | leslielurker wrote:
         | It's not loading for me in Firefox on desktop either; I found
         | the image in the source code if anyone is interested:
         | 
         | https://www.subanima.org/content/images/size/w1200/2021/11/b...
         | 
         | edit: looks intentional?                   /* Remove feature
         | image from top of articles */         .gh-article-image {
         | display: none;         }
        
       | LASR wrote:
       | This concept blew my mind when I internalized it.
       | 
       | Same reason why honest signals exist. A peacock with very rich
       | feathers is a fitness disadvantage. But they find mates more
       | successfully. These traits persist in the gene pool.
       | 
       | It's so much easier to just evolve a cheating trait that does the
       | job of finding a mate even without the required fitness.
       | 
       | But the signals stay honest for the most part.
       | 
       | Why?
       | 
       | It's because ultimately the species survives, not the
       | individuals.
       | 
       | In a lot of cases, something that makes the individual more fit
       | also makes the species more fit. But in some cases, they are
       | inversely proportional.
       | 
       | Hence you end up with suicidal genes that favor the death of the
       | individuals for the greater good of the species.
       | 
       | Now extrapolating to human society, most nations have landed on a
       | system where taxes are paid to the government. Every individual
       | might complain and try to get out of paying. But we do. Why?
       | Maybe because societies where that wasn't a thing were less fit
       | and didn't last long enough to still be around.
        
         | pfdietz wrote:
         | > It's because ultimately the species survives, not the
         | individuals.
         | 
         | No, this is wrong. "Survival of the species" isn't a basis for
         | selection. It will not lead to a gene becoming more prevalent
         | in a population of interbreeding individuals.
         | 
         | Bees sacrifice themselves because they share genes with the
         | queen; genes that are involved in this sacrifice increase their
         | relative abundance in the bee gene pool by increasing the
         | fitness of the superorganism that is the colony.
        
           | Salgat wrote:
           | That's not entirely true. For example, being gay is
           | hypothesized to give an evolutionary advantage because you
           | can provide care for your sibling's children, who share their
           | dna with you. Same goes for early menopause. That can extend
           | to small villages where individuals may give up their own
           | resources for a greater survival chance of their kin within
           | the collective.
        
             | asingnh wrote:
             | Are homosexuality and early menopause genetic conditions?
        
               | SoftTalker wrote:
               | Seems quite likely to me.
        
               | jahewson wrote:
               | Everything that makes us human is constrained by the
               | possibilities offered by our genes. Epigenetics,
               | development, and environment are downstream of that. It
               | is our genes that allow for sexual reproduction in the
               | first place and why we're attracted to other humans and
               | not, say, trees.
        
             | alt227 wrote:
             | Pre 1800, the average life expectancy was aged between
             | 20-40 [1]. I think the menopause is something that was
             | experienced by extremely few people until after then.
             | 
             | [1] - https://en.wikipedia.org/wiki/Life_expectancy
        
               | olddustytrail wrote:
               | Average life expectancy is misleading. You want perhaps
               | median life expectancy after the age of 20.
        
           | meindnoch wrote:
           | >"Survival of the species" isn't a basis for selection. It
           | will not lead to a gene becoming more prevalent in a
           | population of interbreeding individuals.
           | 
           | Well, "species" is but a loosely defined set of genes.
        
             | pfdietz wrote:
             | And group selection cannot increase the frequency of a gene
             | in that collection.
        
         | s1artibartfast wrote:
         | I think you are missing a few points. First, is the adversarial
         | nature of mate selection.
         | 
         | A female peacock who falls for a trick will have fewer
         | offspring that survive. The discerning hen will do better.
         | Honest communication works because it is backed up actual
         | fitness. It doesn't require group selection.
         | 
         | Second, I think there is a lot more going on with respect to
         | taxes. Taxes have existed for maybe 10,000 years. An armed man
         | demanding half your stuff or they kill your family is a tax
         | too. Same for a mature lion that eats what another animal
         | killed. I would argue taxes are an inherent result of power
         | imbalances among humans. Differences give rise to power
         | differentials, which give rise to security concessions, which
         | consolidate into kingdoms and nations.
        
           | notahacker wrote:
           | > Taxes have existed for maybe 10,000 years. An armed man
           | demanding half your stuff or they kill your family is a tax
           | too. Same for a mature lion that eats what another animal
           | killed. I would argue taxes are an inherent result of power
           | imbalances among humans. Differences give rise to power
           | differentials, which give rise to security concessions, which
           | consolidate into kingdoms and nations
           | 
           | Tax fits the model pretty well. Defending against bandits who
           | steal everything and move on is expensive, so kings that
           | claim much smaller portions of wealth and scare off bandits
           | tend to lead to better nations. (Then you've got modern
           | democracies, that typically tax much more, but in a way which
           | is actually compatible with higher growth because the money
           | tends to be spent back into the sluggish parts of the economy
           | rather than spent on zero sum competition with neighbouring
           | kings/lords over territorial tax bases and precious import
           | collection)
        
       | WalterBright wrote:
       | A fascinating read about such things is "The Red Queen" by
       | Ridley.
       | 
       | https://www.amazon.com/Red-Queen-Evolution-Human-Nature/dp/0...
       | 
       | It's all about the propagation of the genes, not the survival of
       | the organism.
        
         | n8henrie wrote:
         | Read this in my early 20s and _loved_ it. Many ideas that have
         | stuck with me. Hoping to reread it with my wife soon, nearly 20
         | years later, and see how it aged.
        
       | WalterBright wrote:
       | Why do humans live long enough to be grandparents? It's because
       | grandparents take care of the grandkids while the parents work.
        
         | derektank wrote:
         | I find the grandparents hypothesis compelling at first glance
         | but it sort of begs the question, why don't we live
         | indefinitely in the first place? There are obvious answers, we
         | make tradeoffs to improve performance early in life at the
         | expense of long term function, but it doesn't seem like the
         | reproductive benefits of caring for a grandchild that only
         | shares a 1/4 of your DNA necessarily tips the scales of
         | selection towards longevity. Especially when, in theory, men
         | remain fertile their entire lives and thus there should have
         | always been some selection for longer life spans. You would
         | expect the reproductive benefits of a 70 year old caring for
         | their own child might be at least comparable to 70 year old
         | caring for a grandchild.
        
           | lcnPylGDnU4H9OF wrote:
           | > why don't we live indefinitely in the first place?
           | 
           | It doesn't seem to be necessary for the survival of our
           | genes.
           | 
           | > You would expect the reproductive benefits of a 70 year old
           | caring for their own child might be at least comparable to 70
           | year old caring for a grandchild.
           | 
           | They're competing with 20-30 year-olds with better physical
           | fitness for a mate. This would be relevant for ~99% of human
           | existence even if it's not totally relevant today.
        
           | WalterBright wrote:
           | why don't we live indefinitely in the first place?
           | 
           | 1. need to make room for the young, ecosystems are not
           | unlimited
           | 
           | 2. we accumulate parasites and diseases as we live. Dying
           | kills them off, too
           | 
           | 3. much slower evolution, implying losing ground compared
           | with quickly evolving competitors
        
           | thrance wrote:
           | Members of some species take care of the children of others,
           | as well as theirs (Orcas come to mind, humans too if course).
           | There is an advantage in helping others with similar DNA than
           | you, because they will reciprocate.
        
           | EdwardDiego wrote:
           | Obviously because Tree-of-Life didn't survive on Earth, so we
           | could never achieve our final form.
           | 
           | https://en.wikipedia.org/wiki/Pak_Protector
        
         | jajko wrote:
         | I don't think its that simple. If we look back far enough, it
         | was more like the man/men hunted or gathered and women took
         | care of kids, fire and cooking.
         | 
         | If I look at less distant ancestors, they all worked in the
         | fields, and so did grandparents (who were not as old as these
         | days when 15 was a good age to start bearing kids, so 35 years
         | old granny was normal). So it again falls mostly on women.
         | Grandparents, those still living, much less.
        
         | hinkley wrote:
         | There's the historian factor as well.
         | 
         | They've found that African Elephant populations are largely
         | constrained by water availability. Creating artificial watering
         | holes is helping restore elephant populations better than most
         | other attempts.
         | 
         | But the matriarch is typically one of the oldest female members
         | of the group, and elephants remember watering holes that they
         | haven't visited since they were young. During a drought they
         | will check all of these secondary and tertiary water sources.
         | If that elephant is killed by poachers, the herd may lose the
         | last remaining record of water resources and suffer for it.
         | 
         | I also recall watching a documentary about a troupe of
         | primates. They adopted a young male kicked out of another
         | troupe. Nothing remarkable about him until, again, a drought
         | year. Turns out not all knowledge of edible foods is
         | instinctual. They discovered him eating a fruit none of the
         | tribe had eaten before. When he didn't die they all started
         | eating it too.
         | 
         | So I think we underestimate the value of record keeping with
         | respect to longevity and inter-group mixing. It's not all genes
         | and safety in numbers.
        
           | EdwardDiego wrote:
           | In Maori culture in NZ, tribal elders (kaumatua) function
           | much the same, they are the keepers of knowledge, history and
           | cultural practices for their group(s).
           | 
           | I suspect you'd find the same in many cultures that didn't
           | have writing to transmit knowledge.
           | 
           | (There's also the aspect of "mana", a very hard word to
           | translate, "face", "influence", "power" or "really respected"
           | are good approximations.
           | 
           | A kaumatua with mana accords your group (iwi/tribe or
           | hapu/sub-tribe) with their mana.
           | 
           | So, say your Granddad was a renowned warrior or diplomat,
           | even though he's in his 80s, the respect his opponents
           | developed for him still persists, and flows onto you.)
        
       | spqr0a1 wrote:
       | While a bee stinger may get stuck in you, that's not so when
       | stinging fellow insects.
       | 
       | The barbs don't catch on an exoskeleton like they do for thick
       | and elastic mammalian skin.
       | 
       | An elegant way to deliver more venom to larger targets.
        
         | randall wrote:
         | Wow that's super interesting! What a novel mechanism.
        
         | wizzwizz4 wrote:
         | If you're careful with the index fingers of opposite hands, you
         | can remove the stinger from your skin without killing the bee.
        
           | shakna wrote:
           | I don't think I've ever been stung in such a convenient
           | position as to allow that.
        
           | 867-5309 wrote:
           | as opposed to index fingers of the same hand..
        
             | qup wrote:
             | Not the same hand, he's talking about two hands on the same
             | side
        
               | mech422 wrote:
               | but I need my second left hand to film for Insta! :-P
        
               | koolba wrote:
               | There's only one frood who could pull off that
               | extraction.
        
           | cryptonector wrote:
           | Not really. The bee that stings you will flap her wings very
           | vigorously, and it will rip its stinger off in less than a
           | second trying to get away from you. Unless you're
           | deliberately trying to get stung and save her, you won't have
           | a chance.
        
             | wizzwizz4 wrote:
             | Depends on the colony. The bees that have stung me have
             | always taken 5 or 10 seconds to start trying to dismantle
             | themselves in earnest, which (depending on location) is
             | usually enough time to rescue them. (I'm not sure whether
             | they _survive_ my rescue, but at the very least they can
             | fly away, and their stingers don 't remain in my skin.)
        
         | EdwardDiego wrote:
         | Yeah, I was hoping the article would mention this, but no dice.
         | :(
        
         | ErigmolCt wrote:
         | Nature's design is often elegant. But also (sometimes) cruel.
        
       | pizzafeelsright wrote:
       | Can someone answer this without an evolutionary presupposition?
        
         | krapp wrote:
         | What are you asking for, a parable? A metaphor?
        
         | gnkyfrg wrote:
         | Dying worker bees ensure survival of the group without a
         | measurable impact on death of the colony, which when seen as a
         | super organism, means only a part of the organism, leaving the
         | reproducing parts intact, since workers don't mate anyway.
        
         | krisoft wrote:
         | It is in the article "A honey bee dies when it stings you
         | because its stinger is covered in barbs, causing its abdomen to
         | get ripped out when it tries to fly away. And surviving with
         | your guts spilling out everywhere is pretty bloody hard."
         | 
         | That's the baseline answer. It is a simple observation, and at
         | this level of question you don't need to worry about evolution
         | at all.
        
         | pestatije wrote:
         | they are so pissed off by your presence that they say fuck it,
         | ill fuck this bastard no matter what
        
       | gnkyfrg wrote:
       | > we humans and nearly all other mammals certainly don't have a
       | singular queen and a sterile caste of workers.
       | 
       | No we have many queens and many sterile workers. Almost all women
       | have children and increasingly more men don't father children.
       | We've outlawed polygamy, but society is trending that way anyway,
       | especially considering online dating statistics. 80% of women are
       | choosing from the top 20% of men.
       | 
       | Men die in war. Look at ukraine. A million men dead, while women
       | dance in clubs. Vietnam, on the American side: 55,000 men dead, 8
       | women dead. Most women stayed home, like the queen bee. Most of
       | those men were sterile for all intents and purposes.
       | 
       | If we don't introduce artificial measures, the natural tendency
       | is toward fewer women mating with more men and the end result is
       | one to many. There's a British lady trying to mate 1,000 men.
       | 
       | We are more like bees than we like to think.
        
       | lysace wrote:
       | A random bee sting _in class_ was the straw that broke my back in
       | a mid 90s multivariable calculus lecture at a Swedish university
       | where I was studying CS /EE. It lead to me dropping out. Went to
       | a local internet/web software startup instead and a whole new
       | world opened up.
       | 
       | Yes, I had been behind. I'm doing OK now :)
        
         | hinkley wrote:
         | As an easily distracted high schooler just trying to enjoy one
         | of his favorite classes, I discovered I could swat a flying bee
         | dead with my folder. They were getting in through some gap in a
         | window facing an alcove an I think we had four or five one year
         | before Facilities fixed the problem.
         | 
         | It worked out, but you don't really want to go squishing bees
         | in an open area since they release chemicals that put their
         | siblings on alert. If they stay put a glass and a piece of
         | stiff paper are a better solution. But these were buzzing
         | around my fellow students making everyone freak out.
        
         | ec109685 wrote:
         | The gate keeping of all that calculus for a CS degree is silly.
         | Wasn't the strongest at math, so grinned and bared but don't
         | really have a grasp of it anymore, and it would have been a
         | shame to not have graduated with a CS degree because of it.
        
           | calvinmorrison wrote:
           | I dropped CS for calc 2
        
             | hightrix wrote:
             | Same. And now I'm 15 years into my software engineering
             | career and the only regret I have is that I didn't spend
             | more time with linear algebra.
        
               | lysace wrote:
               | I know right - that was the fun kind of math.
        
           | lysace wrote:
           | In Sweden it was a heritage from Ericsson. They needed/need
           | engineers who knew that stuff. Supposedly. I should have
           | picked something with less EE even though I also loved
           | electronics.
           | 
           | It seems much better these days.
        
           | cryptonector wrote:
           | It's "grin and bear", as in grinning while bearing the load.
           | The past tense would be "grinned and bore". FYI.
        
         | swyx wrote:
         | basically you are Spiderman
        
           | lysace wrote:
           | Huh. How about that.
        
           | tradertef wrote:
           | beeman
        
       | captn3m0 wrote:
       | As I've been listening to Mythos recently, I must point out that
       | it is also because Zeus cursed the Bee
       | 
       | > In his final response on the matter he declared that she will
       | be a Queen of a colony of workers that will aid her in gathering
       | honey. However, Greek Gods were never truly honourable in their
       | wishes unless it benefitted them directly. In addition to her
       | swarm of workers she was also granted a fatal sting, but this
       | sting would be fatal to her or her colony if they ever used it on
       | another. It was from then on that the honeybees' was barbed;
       | meaning that if their weapon was ever to be "deployed" that the
       | individual that used their sting would not survive the attack.
       | 
       | https://crawliomics.wordpress.com/2019/06/12/zeus-the-honeyb...
        
       | jovial_cavalier wrote:
       | >Thirdly, the haplodiploidy hypothesis only works if all sisters
       | share the same father and if a queen is biased to produce more
       | daughters than sons.
       | 
       | The sex ratio doesn't actually seem like a problem for the
       | theory, because it sounds like for a worker bee, the relatedness
       | of the marginal sibling is 5/8 in expectation, vs. 1/2
       | relatedness of the marginal offspring.
       | 
       | I think you also have to discount the relatedness into the
       | future. If the colony you are born into is already established,
       | your 5/8 related marginal sibling has a much higher likelihood of
       | survival than your 1/2 related marginal offspring when you take
       | into account the risk of breaking from the colony and starting
       | your own.
       | 
       | That probably goes some way to explaining the first problem of
       | multiple fathers. Marginal half-siblings are only 1/4 related to
       | the worker, but they may have a greater chance of survival.
        
       | myflash13 wrote:
       | I don't understand why any "why" question in evolutionary biology
       | is ever satisfied with a "survival of the fittest" truism. Any
       | evolutionary explanation you give me is also falsified by the
       | existence of another species with a different/opposite trait.
       | Whatever explanation you give for a bee's barbed stinger is
       | falsified by the wasp's non-barbed one. Also doesn't answer other
       | questions, such as why didn't bees evolve a type of barbed
       | stinger that doesn't rip their guts out and kill them? Or why do
       | they even need a stinger at all, as many insects don't have one?
       | 
       | Evolutionary explanations like these for "why" things are the way
       | they are not just pure idle speculation, but they are also often
       | unprovable, unfalsifiable, and have the same circular logic as
       | bad religion. Why do species survive? Because they were the
       | fittest, because they survived. But why?
        
         | sedatk wrote:
         | Wasps and bees have different ecological constraints with
         | different risks involved. There is no contradiction here. They
         | evolved as the fittest in their own constraint set. If bees
         | weren't fit enough, they would have gone extinct and replaced
         | by bees with non-barbed stingers. There is no magic that makes
         | them survive.
         | 
         | Evolution doesn't have any goals or agenda. That's why whales
         | still have vestigial hip bones despite having no hips
         | whatsoever. Because it's not a significant parameter in their
         | survival. Same with barbed stingers of bees.
        
           | jstanley wrote:
           | I think this is a perfect example of what your parent comment
           | is talking about, being:
           | 
           | > unprovable, unfalsifiable, and have the same circular logic
           | as bad religion.
           | 
           | You said:
           | 
           | > There is no contradiction here. They evolved as the fittest
           | in their own constraint set. If bees weren't fit enough, they
           | would have gone extinct and replaced by bees with non-barbed
           | stingers. There is no magic that makes them survive.
           | 
           | Sure, there's no _contradiction_ , but this is totally
           | circular reasoning that could be used to prove anything.
           | 
           | The connection graph between "They survive because they're
           | the fittest" and "They must be the fittest because they
           | survive" is just a circle disconnected from all other
           | knowledge.
           | 
           | Your strength as a rationalist is your ability to be more
           | confused by fiction than by reality.[0]
           | 
           | But with this circular understanding of natural selection,
           | you could be given a description of _absolutely any_
           | conceivable configuration of organism and your response would
           | be the same:  "they must be the fittest, because they
           | survive, because only the fittest survive" and you haven't
           | gained any understanding at all.
           | 
           | There will never be a contradiction, because the argument is
           | disconnected from any larger system of reasoning that could
           | plausibly contradict it.
           | 
           | "Hey, there is a random monkey in the Amazon that has 3 hoops
           | on its head and a big hole through its abdomen, isn't that
           | weird? Why are they like that?"
           | 
           | "Ah, the hoops and the holes are required for Fitness. Only
           | the Fittest survive, you know. So if they have 3 hoops on
           | their heads and big holes in their abdomens, that is what
           | makes them Fittest. Amen."
           | 
           | "Why aren't other monkeys like that then?"
           | 
           | "Other monkeys don't need hoops and holes for Fitness.
           | Otherwise they too would have hoops and holes. :)"
           | 
           | A better understanding of natural selection would be
           | _confused_ about the hoops and the holes, and that
           | _confusion_ would correlate with either the random monkey
           | species actually not existing, or the model being wrong.
           | 
           | As regards the bees: there probably is a reason that dying
           | when stinging confers Fitness. But we should _find out_ what
           | that reason is, rather than state  "Fitness because Survival"
           | and feel like we've answered the question.
           | 
           | [0] https://www.lesswrong.com/posts/5JDkW4MYXit2CquLs/your-
           | stren...
        
             | Retric wrote:
             | > totally circular reasoning that could be used to prove
             | anything.
             | 
             | No, you can attack the reasoning by looking into actual
             | costs. It seems like it can explain anything because we
             | don't constantly see examples where it's false.
             | 
             | Looking at the costs to bees you see what percentage of
             | them die from attacking mammal flesh and yep it's a tiny
             | rounding error.
             | 
             | Hypothetically, in a world without constraints mice could
             | have a 100 foot long teeth, but we don't live in a world
             | without constraints.
        
               | xyzzy_plugh wrote:
               | > Hypothetically, in a world without constraints mice
               | could have a 100 foot long teeth
               | 
               | Oh boy, today's the day you learn something new about
               | rodents.
        
               | stavros wrote:
               | What, the fact that their teeth can grow so long that
               | they'll circle around and stab their brains?
        
             | kevinventullo wrote:
             | My understanding from the article and the general theory of
             | Superorganisms is that it's not exactly true that "dying
             | when stinging confers fitness". Rather, dying when stinging
             | is just not a huge penalty when you're talking about non-
             | reproducing members of a colony. So, while it may be a good
             | thing for bees to evolve the ability to survive stinging,
             | the selective pressure is not as large as one might
             | intuitively expect.
             | 
             | Maybe a better title for the post would be something like,
             | "Isn't it weird that bees die when they sting? Shouldn't
             | they have evolved away from that?"
        
               | thaumasiotes wrote:
               | Well, there's a larger problem in the post. The primary
               | reason that a bee dies when it stings you is that you
               | kill the bee. A bee stinging an inanimate hunk of meat is
               | unlikely to die.
               | 
               | But they can die, and yeah, a big part of the reason why
               | is that dying isn't as large of a cost for bees as you
               | might expect from a human perspective.
               | 
               | And looping back, another part is that given the very
               | high risk of being intentionally killed when stinging an
               | enemy who you _want_ to sting, improving the much smaller
               | rate of accidental death isn 't really worth much. But
               | even though it isn't worth much, it's worth something,
               | and work has been done on the project.
        
               | tsimionescu wrote:
               | Per everything I've ever read on this, a bee that has
               | stung meat is no longer able to survive. It will either
               | try to pull itself out and disembowel itself, or it will
               | remain stack and die of hunger. What makes you think a
               | bee that stings, say, a dog that can't swat it will then
               | go on to survive?
        
               | thaumasiotes wrote:
               | Watching a youtube video by a beekeeper demonstrating
               | that a bee stuck in his arm will gently wiggle itself
               | around until it can pull free and fly away. Between that
               | one guy with his live demonstration and his plentiful
               | experience being stung by bees, and you with everything
               | you've ever read, I'm obviously going to stick with him.
               | 
               | You should read better sources.
               | 
               | Here, an example of how bees behave after stinging meat:
               | https://www.youtube.com/shorts/7wH8dYiGbig
               | 
               | Or here: https://www.youtube.com/watch?v=G-C77ujnLZo
               | 
               | A bee that stings meat will pull itself out unharmed, _if
               | you let it_.
        
               | cryptonector wrote:
               | Not at all my experience. My Italian bees are pretty hot
               | though, so that may be why. My Russian bees are much
               | nicer, and I've never been even stung by one of them, so
               | maybe that's it. My hot bees will rip their stinger out
               | within a second, and they emit a very particular
               | frequency that I feel and that lets me know very quickly
               | that I'm in trouble -- to be fair I've only experienced
               | that vibration a handful of times because I've gotten
               | much better at not leaving any exposed skin or any way
               | for them to get in my suit.
               | 
               | All those bee rescue videos you see where the beekeeper
               | doesn't wear a veil are not telling you the whole story.
               | Before they go work on extracting the hive they'll first
               | check that those bees are not hot and angry. If those
               | bees are africanized then they'll use a suit and smoke
               | and they will not bother making or posting a video about
               | it. I.e., those videos are all cherry-picked experiences,
               | all the good ones.
        
               | thaumasiotes wrote:
               | Does any of that support the idea that "a bee that has
               | stung meat is no longer able to survive"?
               | 
               | This is clearly false. Why repeat it?
        
               | cryptonector wrote:
               | The ones that manage to leave the stinger in never
               | survive in my case. Idk what to tell you. That's just my
               | experience. Can they survive? Yes, they can. Does it
               | happen? Yes. Does it happen often? No, I don't think so.
        
               | tsimionescu wrote:
               | Here is an example of a bee disemboweling itself:
               | 
               | https://m.youtube.com/shorts/u9YDf9w6Ps8
               | 
               | Here is another one:
               | 
               | https://m.youtube.com/shorts/fMxZ4vKmOZo
               | 
               | So I've countered your two anecdotes with two anecdotes
               | of my own. Can we go back to looking at what people
               | studying bees have written?
               | 
               | For example, this article interviews two bee scientists
               | that confirm that the majority of honeybees die via self-
               | disembowelment after stinging a human:
               | 
               | https://www.livescience.com/do-bees-die-after-stinging
               | 
               | However, they mention an interesting other thing, that
               | may actually help explain what happens much better: bees
               | don't die when they sting other insects or spiders, they
               | only die when their needles get stuck in our thick skin.
               | So perhaps the most likely explanation for the evolution
               | and survival of the barbed stinger trait is that it's
               | beneficial when bees fight their common enemies, and that
               | bees simply don't interact that much with mammals and
               | their thick skins.
        
               | imtringued wrote:
               | I'm not sure why this answer is buried down so deep. It
               | should be pretty obvious.
               | 
               | The average bee has no reason to ever fight a mammal.
               | Optimizing for a rare event like that makes no sense.
               | It's only once humans started domesticating honey bees
               | that the interaction between honey bees and mammals has
               | become a frequent occurrence.
               | 
               | Surviving attacks against humans that help you survive is
               | a really strange priority.
        
               | tsimionescu wrote:
               | To be fair, many forest mammals will try to eat honey
               | (bears and honey badgers being just two well known
               | examples) and those have skins much thicker than even
               | ours, so I suspect that bees have the same issues when
               | stinging them. And those mammals are much more of a
               | threat to a hive than insects and spiders, as they will
               | virtually destroy the entire hive structure if they are
               | not deterred, and likely kill thousands of larvae and
               | bees.
        
               | thaumasiotes wrote:
               | > It's only once humans started domesticating honey bees
               | that the interaction between honey bees and mammals has
               | become a frequent occurrence.
               | 
               | This can't possibly be true; the honeyguides are a family
               | of birds whose evolved behavior is to find humans and
               | lead them to wild beehives so that the humans can forage
               | the honey (and the honeyguide can forage the leftovers).
        
               | thaumasiotes wrote:
               | > So I've countered your two anecdotes with two anecdotes
               | of my own. Can we go back to looking at what people
               | studying bees have written?
               | 
               | No? You claimed that bees stinging meat cannot survive.
               | Your anecdotes do nothing to support that. Decide what
               | you want to say, then look for support. Where do you get
               | the idea that two examples of something happening provide
               | just as much support for the idea that it always happens
               | as four examples of it not happening do for the idea that
               | it doesn't always happen?
               | 
               | But for what it's worth, Aristotle wrote in the 4th
               | century BC that bees stinging humans often recover, but
               | that they will inevitably die _if they lose their
               | stinger_.
        
               | tsimionescu wrote:
               | This presumes that proto-bees had stingers that killed
               | them before being super organisms, so that the obvious
               | survival disadvantage that dying after a successful
               | attack brings was compensated by the hive life rather
               | than by surviving the sting.
               | 
               | I'm not at all sure this is true - I don't know the
               | evolutionary history of bees, but it seems unlikely that
               | some kind of solitary proto-bees would have died after a
               | sting. And even if this were true, we should still wonder
               | why that proto-bee evolved to have this suicide stinger
               | in the first place.
               | 
               | "It's not a big disadvantage to survival" can't be the
               | explanation for a trait, unless that trait is a remnant
               | from an ancestor where it brought an advantage (like the
               | hip bones in whales - hip bones are obviously useful in
               | land-based mammals, and whales are descendants of those).
               | 
               | Sp the question is: why did some organism ever evolve a
               | stinger that kills it, how was that ever something that
               | made some organism survive better than its brethren that
               | didn't have this trait?
        
               | xyzzy_plugh wrote:
               | > This presumes that proto-bees had stingers that killed
               | them before being super organisms, so that the obvious
               | survival disadvantage that dying after a successful
               | attack brings was compensated by the hive life rather
               | than by surviving the sting.
               | 
               | I don't see how you arrived at this conclusion, this
               | logic seems to be flawed.
               | 
               | Of course it can be an explanation for a trait. If you
               | are genetically predisposed to high cholesterol, is that
               | advantageous or disadvantageous? There's simply too
               | little pressure to do anything about it.
               | 
               | Phenotypes aren't required to change in the smallest
               | imaginable step. It's not implausible that nature decided
               | "hey this next bee gets some furry yellow stripes, but
               | also barbs" and here we are.
               | 
               | Not everything is optimal in the extreme. For all we know
               | there have been many, many bees without barbs, but the
               | bar to pass that on as an advantage is very high. The
               | odds of a bee reproducing aren't even that high to begin
               | with.
        
               | tsimionescu wrote:
               | > Of course it can be an explanation for a trait. If you
               | are genetically predisposed to high cholesterol, is that
               | advantageous or disadvantageous? There's simply too
               | little pressure to do anything about it.
               | 
               | If barbed stingers were a variation that occurred in some
               | individual bees, but many other bees didn't have them,
               | like predisposition to cholesterol in humans, then I'd
               | agree.
               | 
               | But a trait can't be universal in a species, and even in
               | many related species, unless there is explicit selective
               | pressure _for_ that trait, or if it 's a remnant from a
               | common ancestor that had it. Your high cholesterol
               | predisposition example is actually perfect, it shows what
               | happens with traits that don't have significant pressure
               | for or against them: they remain confined to a subset of
               | the population.
               | 
               | > Phenotypes aren't required to change in the smallest
               | imaginable step. It's not implausible that nature decided
               | "hey this next bee gets some furry yellow stripes, but
               | also barbs" and here we are.
               | 
               | They generally are. The likelihood of a random mutation
               | that produces several phenotypal changes but leaves the
               | organism still viable is extremely low.
        
               | xyzzy_plugh wrote:
               | > If barbed stingers were a variation that occurred in
               | some individual bees, but many other bees didn't have
               | them, like predisposition to cholesterol in humans, then
               | I'd agree.
               | 
               | What makes you believe they are _not_ a  "variation" in
               | individual bees? Colony genetics are fascinating and can
               | facilitate some pretty extreme variation.
               | 
               | Could a single queen's mutation be responsible for all
               | barbs? You have some hidden constraints here that I don't
               | think we agree on.
               | 
               | > They generally are. The likelihood of a random mutation
               | that produces several phenotypal changes but leaves the
               | organism still viable is extremely low.
               | 
               | This is simply wrong. There are always plenty of
               | potential mutations that cripple viability, sure. But
               | pleiotropy in general is relatively common. In some
               | species more than others.
               | 
               | Your reasoning here is heavily flawed. You firmly believe
               | evolution and genetics work in a particular way that
               | simply does not reflect reality. You are putting your
               | belief in "survival of the fittest" first. I encourage
               | you to broaden your horizons.
        
               | heavyset_go wrote:
               | > _" It's not a big disadvantage to survival" can't be
               | the explanation for a trait_
               | 
               | Why not? Honest question.
               | 
               | Intuitively, I'd say that there are plenty of traits that
               | propagate as long as they aren't expensive in terms of
               | genetic survival. It's the nature of random genetic
               | mutation, random traits will develop and some of them
               | won't really affect survival and may propagate.
        
               | tsimionescu wrote:
               | Yes, traits can propagate even if there is no selective
               | pressure for them. But they won't reach 100% of the
               | species, even less so 100% of several geographically
               | separated species (such as African and European bees)
               | just because they aren't that bad. Could we contrive a
               | story where it could be possible something like this does
               | happen? Sure, but it would be very unlikely (basically,
               | it would require only a small population that happened to
               | have this minor handicap to have randomly survived some
               | mass extinction event that killed off the entirety of the
               | rest of the species).
        
               | mibsl wrote:
               | IIRC genetic drift can reach 100%.
        
               | kevinventullo wrote:
               | _This presumes that proto-bees had stingers that killed
               | them before being super organisms_
               | 
               | I don't know where you got that. It seems perfectly
               | plausible that the stingers evolved in the superorganism,
               | and that the selective pressure was something like
               | "drones protect the hive via stingers which kill them"
               | versus "drones can't sting at all".
        
               | tsimionescu wrote:
               | This is a different argument, one which I can agree with.
               | I was specifically saying that the argument for how a
               | species acquired a universal trait can't be "because it
               | didn't hurt their fitness that much", you have to have
               | positive pressure for a trait to spread to the entire
               | species (or it must be a trait left over from an ancestor
               | where it had these pressures).
        
             | iwontberude wrote:
             | How is it circular to argue why one species would do better
             | in an environment than another based on phenotype and the
             | physical interactions it enables? It's all relative to
             | other species. As long as you understand that, there is no
             | logical fallacy. I do very much appreciate the focus on
             | informal logic though.
        
               | jstanley wrote:
               | Because you could encounter _absolutely any_ organism and
               | make the same argument. There is no configuration of
               | organism that would cause you to say  "huh, I guess
               | Survival doesn't depend on Fitness after all!"
               | 
               | Because it takes the observation of Survival and uses it
               | to infer Fitness, at the same time as saying that Fitness
               | confers Survival.
        
               | glenstein wrote:
               | >Because you could encounter absolutely any organism and
               | make the same argument.
               | 
               | That's a function of the explanation being an extremely
               | good explanation. It rises to the top precisely because
               | it has explanatory power all across nature without
               | evident counter-example.
               | 
               | >There is no configuration of organism that would cause
               | you to say "huh, I guess Survival doesn't depend on
               | Fitness after all!"
               | 
               | This is where the argument falls apart. For starters,
               | species go extinct all the time for reasons tied to their
               | evolutionary trajectory. And there are species still
               | living that unfortunately seem very imperfectly adapted
               | to their constraints and likely to go extinct without a
               | run of good luck or human intervention (e.g. pandas). We
               | seem perfectly capable of recognizing when such species
               | are "on the ropes". Additionally there are relative
               | advantages we can clearly observe from animals in
               | overlapping niches, and we can marvel at the
               | effectiveness of adaptations in ways that don't involve
               | circular assumptions (e.g. algae's capability for
               | efficient growth is astonishing and without equal on the
               | planet).
               | 
               | And, we could surely conceive of preposterous examples
               | that defy expectations (e.g. the other commenter's
               | example of mice with 100ft teeth).
               | 
               | It probably _feels like_ it proves too much, because it
               | 's confirmed over and over again in nature everywhere, at
               | all times. But in an alternate world where that wasn't
               | the case, counter-examples would abound (such as the
               | mouse with 100ft teeth). So re-iterating the core lesson
               | about the role of natural selection is not just a
               | circular assumption, it's the culmination of hard earned,
               | accumulated evidence, ready at any moment to be
               | falsified.
               | 
               | The honeybee is a perfect example, because the stinger
               | _does_ pose a real question about how we understand it 's
               | relation to fitness, and it requires delving into all
               | kinds of complicated dynamics about genetically related
               | drones are to the queen, the role of the sacrifice in
               | supporting the hive and so-on. If we didn't have
               | explanations like those, it would indeed pose a problem
               | with explanations that presume fitness.
               | 
               | That's a real payoff from being alert to the need to have
               | robust explanations; I don't think anyone is just saying
               | "well it's fitness" and calling it a day so much as
               | they're honoring the explanatory power of a well
               | confirmed theory.
        
               | dwattttt wrote:
               | Translating the word Fitness from a term of art makes
               | this very clear: if you said "good enough to survive", no
               | one would question the statement "I wonder why they
               | survive. Guess they must be 'good enough to survive'".
        
               | myflash13 wrote:
               | > It probably feels like it proves too much, because it's
               | confirmed over and over again in nature everywhere, at
               | all times.
               | 
               | No, logically it is proved true because it is assumed to
               | be true and then used to prove itself.
               | 
               | > For starters, species go extinct all the time for
               | reasons tied to their evolutionary trajectory.
               | 
               | Again, this is circular logic. You assumed that the only
               | reason that species go extinct is because it wasn't fit
               | enough. If you assume survival of the fittest then of
               | course it is true.
               | 
               | Here's another circular explanation: things are the way
               | they are because God created it that way. This
               | explanation rises to the top precisely because it has
               | explanatory power all across nature without evident
               | counter-example, right?
        
               | labster wrote:
               | Actually your example of creationist species isn't
               | circular at all, it just has no predictive power. Unless
               | you want to say that God really likes beetles, I suppose.
               | 
               | In the end evolution is random, but exerts some pressure
               | towards fitness in some environments. Some traits are
               | legacy or are just plain random; just because an organism
               | has a trait does not mean it is useful now, or indeed has
               | ever been useful for fitness. The whole package must be
               | reasonably fit for some environment, but that doesn't
               | mean all the traits improve fitness.
        
               | 8note wrote:
               | no, "god created it this way" does not answer for
               | extinctions. if god created it that way, the species
               | would not be extinct.
               | 
               | the part i think youre missing is that "survival of the
               | fitness" is shown elsewhere, and used as a tool here to
               | identify what the fitness is, and how and when certain
               | traits were beneficial.
               | 
               | the case you are descibing is that all applicatioms of
               | science(well, of anything) are circular reasoning. if you
               | use newton's mechanics to predict motion of a mass
               | undergoing acceleration, its circular because your result
               | is proof of newtons mechanics, and newtons mechanics is
               | proof of your result.
               | 
               | its just an "if and only if" relationship. that's not
               | circular reasoning.
        
               | labster wrote:
               | You're assuming God didn't get tired of having the
               | species around so He decided to do some exterior
               | redecorating in His great wisdom.
               | 
               | Also you don't understand physics, the proof is that you
               | can make predictions and then verify the results.
        
               | glenstein wrote:
               | >Again, this is circular logic. You assumed that the only
               | reason that species go extinct is because it wasn't fit
               | enough.
               | 
               | Again, I gave numerous examples, and you ignored mostly
               | all of them.
               | 
               | Just to develop the point about extinctions a bit
               | further. We know for instance of birds that had no
               | natural fear of predators on the Galapagos Islands that
               | upon human contact were driven to extinction or near
               | extinction, and we don't need any circular assumptions to
               | tie their fitness to their extinction.
               | 
               | And that's not to mention the rest of the examples I
               | listed that you ignored, such as cases where we can
               | observe that currently living species are doing well or
               | poorly, like algae and pandas respectively, all of which
               | hinge on knowledge of specific biological mechanisms.
               | 
               | So I don't know why you keep asserting that it's an
               | assumption.
        
               | thowawatp302 wrote:
               | This whole line of argument from the parent comment and
               | its ancestor seems to ignore we can see where certain
               | evolutionary features reach local minima, like the retina
               | of the human eye. It's obviously not the fittest thing
               | wrt the octopus retina, but does well enough you can read
               | this.
        
             | notRobot wrote:
             | "Fittest" is what we call those who happen to survive in
             | their context. Systems that successfully replicate
             | themselves in their context tend to stick around. Those who
             | can't, _go extinct_. We obviously still study why they
             | survived. That 's what the article speculates about. So
             | yes, in a sense, any organisms you see is the "fittest" in
             | the sense that it was able to survive (replicate) in its
             | context while countless others were not.
        
             | icehawk wrote:
             | > The connection graph between "They survive because
             | they're the fittest" and "They must be the fittest because
             | they survive" is just a circle disconnected from all other
             | knowledge.
             | 
             | The GP said that bees survived because they're "fit enough"
             | not that they survive because they're "the fittest" and
             | there are definitely species that don't seem to be
             | surviving because they're not fit enough.
        
             | shwouchk wrote:
             | All of evolution is path dependent.
             | 
             | Dying after a sting does not have to confer extra fitness
             | to exist, right now. rather it had to have conferred some
             | fitness relative to the alternative traits circulating at
             | the time it was selected. obviously if you go by gradient
             | descent you are not guaranteed to reach a global minimum or
             | even a local one, given a constantly changing fitness
             | landscape.
             | 
             | In most of these discussions, optimizing nature of
             | evolution is taken as granted - we do not need to prove how
             | evolution works yet again - there are plenty of evidence
             | and discussion elsewhere - take it or leave it.
             | 
             | This node is well connected to other knowledge, and if you
             | disagree, you need to convince a whole discipline of
             | science, not me.
             | 
             | From the optimizing premise of evolution, various inferred
             | hypotheses can be made, explaining a range of phenomena,
             | just like, in physics, from the premise that probabilities
             | of events are given by the amplitudes of solutions of
             | certain pdes with specific initial conditions, we were able
             | to devise tractable mathematical models of various nuclear
             | reactions, here a model of development of certain abstract
             | traits was explained ("altruism").
             | 
             | The author fully acknowledged that this is a simplified
             | model and does not match reality in some cases, and in
             | other cases does not explain well enough. improvements to
             | the model were proposed.
             | 
             | Isn't that how science works, in the best cases?
        
             | darkerside wrote:
             | This assumes that Fitness is a meaningless aphorism. I
             | would posit that Fitness is a meaningful concept that can
             | be learned. It is defined by what happens to be
             | reproductively helpful to a species, which is tautological,
             | but to understand the definition, it just means you need to
             | understand that particular species ecosystem and lifecycle.
             | 
             | If you really analyze any word, it loses all meaning except
             | what we've assigned it.
        
         | RangerScience wrote:
         | Survival of the fittest is the flawed quote, usually used by
         | those with supremacist conceptual frameworks (that there can be
         | an objective "better", etc). This shows up a lot in fiction,
         | where the quote is used as justification for cruelty,
         | atrocities, and the like.
         | 
         | IMHO, the much better quote is:
         | 
         | > It is not the strongest of the species that survives, nor the
         | most intelligent that survives. It is the one that is most
         | adaptable to change.
         | 
         | See https://www.darwinproject.ac.uk/people/about-darwin/six-
         | thin...
        
           | HPsquared wrote:
           | I'm very tickled by the lack of attention to detail here. The
           | article does present that quote, yes, but it's preceded by
           | the sentence:
           | 
           | "None of the fake soundbites is more insidious than the
           | first:"
           | 
           | i.e. it's a fake quote
        
           | ch4s3 wrote:
           | > It is the one that is most adaptable to change
           | 
           | That is what fit means.
        
           | mixmastamyk wrote:
           | It is not only the best adapted class that survives, aka the
           | "fittest." They only need to be good enough to survive and
           | reproduce. In other words, the principle should be stated as
           | "survival of the good enough." I know it doesn't roll off the
           | tongue as well, but is more accurate.
           | 
           | Perhaps, survival of the fit. (period)
        
         | notRobot wrote:
         | Here's an explanation of how this works:
         | 
         | All creatures are very complicated. Thus reproduction doesn't
         | produce perfect clones, "mutations" take place. This is largely
         | because there are so many different ways to derive one
         | individual out of two individuals' complex genitic material.
         | This is all a feature. This is why individuals have unique
         | characteristics. Think about how different humans are from each
         | other, even though we're all _humans_. This same thing applies
         | to all creatures. Every individual is different. Those who have
         | "disabilities" (disadvantages in their context) are less likely
         | to survive. So those with advantageous traits survive and pass
         | their traits on through reproduction, making those specific
         | traits more prevalent.
         | 
         | The answer to "why didn't x evolve to do y?" is usually just
         | that that specific mutation might have never occurred or caught
         | on randomly. This is also why different species do different
         | things differently. It's all random mutations. Some were
         | beneficial in their context and environment so those who had
         | them were more likely to survive and pass those traits along.
         | 
         | It's not that "the objective of life is to survive" in a
         | spiritual sense, it's that life randomly happens and some of it
         | survives and it makes more life like itself. In some ways, I
         | suppose the purpose of life is to create more life. Systems
         | that replicate themselves successfully survive. We call these
         | "life". It's really a linguistics thing.
         | 
         | Hope some of this makes sense. I enjoyed thinking about this.
        
           | myflash13 wrote:
           | If your answer is "it's a random mutation" then that settles
           | the "why" question permanently. Why all this idle speculation
           | about bee's stingers, then? It was a random mutation, and it
           | survived, done.
        
             | notRobot wrote:
             | It was random, _and it survived_.
             | 
             | Every single part of an organism goes through a
             | recombination/mutation process countless times, the stinger
             | evolved to be what it is today over a very long time and
             | it's cool to study why it ended up the way it did. Tells us
             | about their environment and history and evolutionary
             | pressures, survival is a result of the random traits being
             | successful in their _context_ in specific ways.
        
               | myflash13 wrote:
               | Still doesn't explain why other species in the same
               | context survived without it or with an opposite trait.
        
               | afavour wrote:
               | Sure? Doesn't mean the species-specific examination isn't
               | interesting.
        
               | autoexec wrote:
               | Why shouldn't different or even "opposite" traits also be
               | successful? When faced with random inheritable
               | differences across different species over long periods of
               | time why wouldn't the result be a variety of them, every
               | one of which just didn't prevent reproduction from
               | passing those traits on to the next generation? Some
               | traits might be seen as "better" or "worse" by comparison
               | but as long as they get passed on, we'll see both. It
               | isn't about being "best". It's about being "good enough"
        
               | asdff wrote:
               | Because the trait is not the end all be all. It's a
               | random walk to an outcome that leads to enough offspring
               | to survive for the species to be there at all. It's
               | probably not even the most optimal solution whatever is
               | there, just happens to be competitive among the rest of
               | the ecology to not be snuffed out.
               | 
               | You might be interested in this concept:
               | 
               | https://en.wikipedia.org/wiki/Fitness_landscape
        
         | stouset wrote:
         | > Why do species survive? Because they did, and because the
         | objective of life is to survive. But why?
         | 
         | In evolutionary biology, that _definitionally_ is the ultimate
         | answer. One species survived, another didn't. Sometimes that's
         | because the adaptation helped them outcompete, sometimes it's
         | because they were already competitive and this preexisting
         | disadvantage from an earlier round didn't hurt enough to
         | matter. We can try to find intuitive explanations past that
         | which feel satisfying but it's always going to be a rough
         | approximation.
         | 
         | Let's use chess as an analogy. Allow an engine to analyze a
         | position and tell us the best path forward. But why did it
         | choose that line? We can (and do) come up with explanations
         | that help us fit a move into our understanding of the game:
         | moving this pawn allows that knight to occupy a better spot
         | where it can exert its influence on the rest of the board, or
         | whatever. But that's merely a convenient simplification for our
         | gut understanding. It's not _really_ the actual answer. The
         | ultimate "why" is "because it produces the best possible
         | eventual outcome no matter the response".
        
         | johndhi wrote:
         | Love this comment. It highlights a major misunderstanding of
         | biology that many people who didn't study it in depth have:
         | that every, or most features of living beings do not have an
         | "evolutionary explanation." T-rex arms aren't short so they can
         | open flowers - they just happen to be small because that's the
         | type of creature that happened to survive after a lot of
         | changes.
        
           | bornfreddy wrote:
           | Maybe this explains why humans are snoring? It just wasn't /
           | isn't evolutionary important.
        
             | prerok wrote:
             | Well, the explanation I heard is that snoring provided
             | protection during sleeping to scare away predators. I don't
             | know the source for this theory, so take it with a grain of
             | salt :)
        
               | bornfreddy wrote:
               | I heard that too, but it doesn't sound likely. If I was
               | sleeping and there was a predator passing by, I would
               | prefer it didn't notice me... :)
        
               | pandemic_region wrote:
               | The fact that heavier (and thus more attractive as prey)
               | people are more likely to snore could give credibility to
               | your explanation.
        
             | newsuser wrote:
             | Most likely, yes, like the loudness of baby crying. Humans
             | are pack animals so any predator attracted by snoring or
             | baby cry, and deciding to check it out would be in a very
             | very big trouble.
        
           | dennis_jeeves2 wrote:
           | >many people who didn't study it in depth have
           | 
           | Or are incapable of studying/reflecting in depth.
        
         | pks016 wrote:
         | What kind of explanations are you looking for? Your idea is
         | that there should be some sort of common explanations of why.
         | 
         | I guess because these are theories and best guess based on the
         | evidence. There are many unknowns but that doesn't mean we
         | should disregard what we know.
        
           | myflash13 wrote:
           | If someone simply asked what are the advantages of bees
           | barbed stingers over wasps non barbed ones, that would be an
           | interesting question. But if someone asks "why" and then
           | proceeds to give a circular logic explanation (it survived
           | because it is fit because it survived) that is unprovable, I
           | find that to be silly idle speculation.
        
             | barbazoo wrote:
             | I believe you'd have to look at the evolutionary advantage
             | of bees with barbed stingers vs bees without barbed
             | stingers and how one made that particular group of bees
             | more successful than the other.
        
         | trgn wrote:
         | women go to the bathroom together because if you squat to pee
         | in the tall grass of the savannah you need somebody to lookout
         | for predators
        
         | amelius wrote:
         | The main problem I see with how some popular science
         | journalists approach evolutionary biology is that they always
         | think from the perspective of the individual, as opposed to the
         | group.
        
         | ripitout wrote:
         | Love how the top comment is "I don't see how evolution explains
         | everything in biology". Classic HN.
        
         | crystal_revenge wrote:
         | The "why" questions people ask about evolutionary biology are
         | the carry over of theology into the understanding of evolution.
         | People still need to believe there is a fundamental reason the
         | world is the way it is. A similar theological carry over is the
         | belief that we are better suited to the environment we evolved
         | in. This is akin to "golden age" thinking, that the world today
         | is somehow not right and if we return to the origin things will
         | be better.
         | 
         | At a fundamental level causality doesn't even really make sense
         | in evolutionary biology. You _can_ ask the question  "what
         | benefits do this feature provide", but you can never really say
         | that's _why_ they evolved. In the end you have the traits you
         | do because, at point in the species development, they didn 't
         | make you die faster and some helped you survive better, but
         | it's not really possible to disentangle these.
         | 
         | Likewise people don't really understand that in evolutionary
         | processes both the species _and the environment_ are constantly
         | changing. The notion that a species is  "adapted for a
         | particular environment" is somewhat nonsensical because "the
         | environment" is never really _fixed_.
        
           | atorodius wrote:
           | > The notion that a species is "adapted for a particular
           | environment" is somewhat nonsensical because "the
           | environment" is never really fixed.
           | 
           | Mever considered this. Good stuff
        
         | wruza wrote:
         | Imagine taking your favorite fractal <formula here> and picking
         | a random point in one of its non-trivial regions, trying to
         | explain what happens there. Would you be better satisfied by
         | <formula> or by specific step by step calculations that lead to
         | that neighborhood?
         | 
         | Either way, now imagine taking not your favorite, irregular,
         | non-describable, non-computable, enormously complex processes-
         | driven fractal that is the real nature, then picking a random
         | point in one of _its_ non-trivial regions, trying to explain
         | what happens there. Now ask yourself the same question and what
         | comes to mind.
         | 
         | More short analogy would be that biology is physics with all
         | elementary particles being different.
        
           | enugu wrote:
           | This is an interesting analogy, thanks.
           | 
           | But, doesn't this match the parent comment's point? If
           | someone asks why a particular point and not, a nearby point
           | is on the fractal, the reply is just that complicated program
           | generated this number and not another number. It leads to the
           | theme of incompressibility (is there a program of smaller
           | length which can correctly check whether a point in a given
           | region is on the fractal). Sometimes, programs are
           | incompressible, and at other times there is a local
           | explanation.
           | 
           | So, people writing on this topic should be careful to make
           | this distinction and check if their local explanation (which
           | is a 'compression' of the evolution process) actually
           | classifies points into fit/unfit regions with the assumption
           | that the points are in some common region(shared features in
           | the environment).
           | 
           | Compare also with explanations of historical events(list of
           | causes for a victory but with the same causes present, there
           | is failure in another context) and atheist - religious
           | debates on God's grace for a particular prayer and allowing
           | the suffering of others. If a local explanation can't
           | distinguish events, then the global explanation has to be
           | validated by something else.
        
             | wruza wrote:
             | This analogy doesn't really answer it, it only shows that
             | there's no generic answer. So "fittest" basically means
             | "the result". It's a definition, not explanation. But it is
             | used instead of "the result" because we have some insights
             | into _how_ it was calculated, and we know that the nature
             | simply creates {mi,bi,tri}llions of variants and then some
             | minuscle amount has better chances to pass and mix these
             | genes, outplaying others in this game. That's where fittest
             | comes from. Any attempt to explain it fully meets with a
             | local complexity, in which only high-level questions can be
             | answered in the same limited high-level manner. So there is
             | some compressibility, but it's clearly jpeg. Cause if a
             | whole planet of distinct particles would strictly follow
             | some rule to compress the observations into, it would be a
             | rule of physics, not biology. E.g. why do animals fall to
             | the ground? Due to gravity, here's  <formula>, works for
             | all particles.
        
               | enugu wrote:
               | Sure, you won't get exact predictions on which genetic
               | codes are realized. Similar to how statistical physics
               | doesn't predict microstates, but the macro picture is
               | understood. Even that might be too ambitious for
               | complicated systems, but you can still have sensible
               | predictions from evolution theory which looks at a
               | landscape and says that there will be animals with tall
               | necks and predators with very fast speeds even if you
               | didn't know beforehand about giraffes and cheetahs.
               | 
               | In the fractal analogy, the compressed theory does not
               | predict that exactly this point does/does-not lie on the
               | fractal, but predicting whether a small disc around the
               | point intersects the fractal.
        
         | raincole wrote:
         | I think you misunderstood what people mean by "why" in the
         | context of evolution.
         | 
         | For example, you ask a random person what his job is.
         | 
         | He: I fix TVs
         | 
         | You: Why?
         | 
         | He: Uh, that's what keeps a roof over me and keeps my family
         | fed?
         | 
         | You: But clearly other humans do other jobs and still have
         | roofs. So it's not a real "why". Your statement is falsified.
         | 
         | > Evolutionary explanations like these for "why" things are the
         | way they are not just pure idle speculation, but they are also
         | often unprovable, unfalsifiable, and have the same circular
         | logic as bad religion
         | 
         | How DNA works at molecular level is science. How creators
         | became what they are now is _history_. History usually doesn 't
         | have the same level of falsifiability as science does.
        
           | raincole wrote:
           | * creators -> creatures
           | 
           | This typo somehow fits the theme in a discussion over
           | evolution ;P
        
         | kragen wrote:
         | > _Any evolutionary explanation you give me is also falsified
         | by the existence of another species with a different /opposite
         | trait. Whatever explanation you give for a bee's barbed stinger
         | is falsified by the wasp's non-barbed one._
         | 
         | The explanation in the article does not reduce to a "survival
         | of the fittest" truism and is not falsified by this example.
         | The article explains at great length why that is, specifically
         | referring to that example.
        
         | _orz_ wrote:
         | I had a very similar feeling until I took a course with one of
         | the leading researchers in the field of protein folding. Two
         | things that he repeatedly mentioned stuck with me a lot:
         | 
         | Evolution is not the survival of the fittest but the not-dying
         | of the unfit. That explains why we have so many different
         | species in the same ecological niche. The example he used was
         | different types of grass on the same field. All of those were
         | fit enough to not die.
         | 
         | The second thing he always repeated was that biology only
         | observes what does or at some point did work. That leads to a
         | huge confirmation bias that research needs to be aware of. Two
         | species might be very similar but just across different sides
         | of the boundary of survival.
        
         | throwaway519 wrote:
         | Why did the spaghetti monster give bee srings barbs?
         | 
         | Because the bees prayed for protection and got saucy about it.
         | 
         | Is that better? Are you really looking for equivalence? Is that
         | fit enough?
         | 
         | Instead of spreading evolution FUD, there are many0laces you
         | could educate yourself. S9meone with a HN acvount is more than
         | able to seek these out. Therefore your motives for this comment
         | seen more disingenuous.
        
         | nejsjsjsbsb wrote:
         | I have a personal theory that genetics and survival just need
         | to be good enough. They may find other local maxima and improve
         | things. But they don't need to be perfect in every dimension.
         | There is room for stuff that doesn't make sense to human logic.
         | In addition because species don't generally breed with other
         | species you need to find those optimisations through chance
         | within a species. And genes are not a feature list. Turn one
         | gene on and something else may turn off. There are trade off
         | too.
        
         | dalmo3 wrote:
         | I can't believe no one has mentioned, this is exactly the
         | teleological fallacy.
         | 
         | I was once very curious about evolutionary science until I
         | realised exactly what you said. It's everywhere. Once you see
         | it you can't unsee it.
         | 
         | If you're going to go that route, at least have the sense to
         | use the word "how", not, "why" and never "because". There's
         | nothing causal in evolution.
        
         | ErigmolCt wrote:
         | Evolution isn't a perfect engineer
        
       | yieldcrv wrote:
       | this is wild because the article starts off with explaining how
       | "why" is a bad question, and then doubles down on this entire
       | thesis explaining why a bee dies when it stings _you_ , a human,
       | and the evolutionary nature supporting it
       | 
       | except, its entirely wrong and the foreshadowing about "why" was
       | super important: bees don't 'expect' to die when they sting you.
       | they can sting many creatures and not get their abdomen ripped
       | out because the barb doesn't get stuck in the thing they stung.
       | _just like wasps_.
       | 
       | so this 20 page dissertation is completely baseless.
        
       | metalman wrote:
       | Bee's are a product of many millions of years of evolution, so
       | the why, is that it works! Ever watch bees?, bugs?, other things,
       | up close and in the danger zone? I do. A bee's stinger will embed
       | in you, or me, and then the venom sack rips right out of the bee
       | and it is possible to watch the venom sack pump venom without the
       | bee itself attached anymore. I was attending my mother in her
       | herb garden and commented on there bieng honey bees around, which
       | she disputed, so I caught one, and held its legs while it tried
       | to sting me, and showed her this, but my hands are so callused,
       | that its stinger would not go in, then I let it go. She says I am
       | an improbable creature, and describes me bieng a half Vulcan and
       | half Klingon.
        
       | jpeloquin wrote:
       | The concept of indirect fitness must be more complicated than
       | explained here. The article explains it as a worker bee sharing
       | 75% of her genes with her sisters, but only 50% with a child, so
       | there is selection pressure for workers to be sterile and self-
       | sacrificing. But few genes actually differ between individuals,
       | so the percentages are much higher. E.g., I share ~ 99% of my
       | genes with each one of you reading this. Assuming honey bees'
       | genetic variation is not much more extreme than human variation,
       | we're talking about 99.5% vs. 99.75% sharing, which sounds more
       | like an explanation of why altruism would be preferred in general
       | rather than uniquely affecting bees.
       | 
       | The article does eventually circle around to acknowledge this,
       | but it's easy to miss and very underdeveloped compared to the
       | discussion of kin selection: "So why do bees die when they sting
       | you? Perhaps because they're disposable parts of a larger super-
       | organism which has evolved by multi-level selection."
        
         | prerok wrote:
         | Hmm, I understand this difference in genes differently.
         | 
         | You and I probably share 99% of effective genes, but still the
         | difference in genes is much greater because there we are
         | comparing the entire DNA. There is a lot of non-affecting DNA.
         | And that is what they analyze when comparing DNA of two
         | individuals in forensics.
        
           | jpeloquin wrote:
           | Based on the information I found, the % difference between
           | two randoms humans in terms of base pairs (including non-
           | coding DNA) is even less than the difference in terms of
           | genes, so the distinction does not materially alter the
           | discussion. Also the article framed its explanation in terms
           | of genes, not base pair sequence.
           | 
           | "Between any two humans, the amount of genetic variation--
           | biochemical individuality--is about .1 percent."
           | https://www.ncbi.nlm.nih.gov/books/NBK20363/
           | 
           | https://book.bionumbers.org/how-genetically-similar-are-
           | two-...
           | 
           | Forensic comparisons are mostly about comparing the number of
           | short tandem repeats at handful of loci, a very small part of
           | the the whole genome.
           | 
           | If you have any information that indicates the DNA similarity
           | between people is less than 98-99% I would love to hear it. I
           | have not personally analyzed the sequences from the 1000
           | genome project to check, and am relying on summaries written
           | by other people.
        
             | prerok wrote:
             | I see, thank you for the explanation!
        
         | GuB-42 wrote:
         | It doesn't matter how much bees have in common. The idea is
         | that in bees, altruistic traits, that is those that produce
         | more sisters by helping the queen have a 75% chance of being
         | passed, because sisters share 75% of a worker bee genes. Most
         | of the genes are the same, of course, bees won't become dogs or
         | anything like that, but a few of them differ, and these are the
         | one that matter.
         | 
         | Could worker bees be fertile and have a selfish traits that let
         | them have more children, they would only have a 50% chance of
         | passing these, because children share 50% of genes.
         | 
         | So: 75% of altruistic genes pass vs 50% of selfish genes.
         | Altruistic genes win. Humans can't pass 75% of their genome
         | this way, so that altruistic genes have no intrinsic advantage
         | over selfish genes.
        
           | jpeloquin wrote:
           | Right, "sharing" here must mean DNA that was cloned from the
           | same ancestral DNA strand, not merely that it shares the same
           | informational content. I got lost in the analogies that frame
           | things in terms of what's "better" for the organism and lost
           | sight of this.
           | 
           | The most important thing from the perspective of replication
           | of a DNA strand is the number of copies of DNA passed to the
           | next generation, and future generations, right? Which would
           | be 0.75 * (mean marginal increase in next-generation sisters)
           | + 0.5 * (mean # offspring). The probability that these next-
           | generation individuals actually get to reproduce in turn
           | would also factor in somewhere.
           | 
           | What's also interesting is that if we take the point of view
           | of the queen (through whom the altruistic genes must pass),
           | the queen's reproductive strategy is relatively few children
           | + hordes of sterile helpers + killing her own sisters. So are
           | we talking about a fitness advantage of altruistic traits
           | (maximizing # sisters), or a fitness advantage from selfish
           | traits [maximizing P(fertile child survival) I guess, since #
           | children is small] that produce hordes of sterile helpers?
           | 
           | Edit: Circling back to the organism perspective, in the sense
           | of "I would gladly give up my life for two brothers or eight
           | cousins.", how many bees is it worth giving up one's own life
           | for in that specific sense? We do have a common ancestor
           | after all and thus a non-zero R-factor.
        
       | erikig wrote:
       | TLDR;
       | 
       | Suicidal altruism and costly signaling for the survival of the
       | super-organism. Also, zombie poison delivery pumps.
        
       | harimau777 wrote:
       | Am I the only one who grew up with bees dying after stinging
       | carrying a sort of unspoken significance or meaning?
       | 
       | I don't think I ever heard someone actually state it, but growing
       | up I had the feeling that bees dying when they sting you was in
       | some sense "significant" because it meant that bees had to be
       | selective in when they chose to resort to violence.
       | 
       | It was almost like an unspoken fable or illustration about the
       | importance of controlling aggression.
        
         | ianbicking wrote:
         | It points to another evolutionary pressure that isn't mentioned
         | as often: if an animal is too aggressive humans will
         | exterminate it.
        
           | harimau777 wrote:
           | I had this experience with a wasp nest near my house. I
           | figured "live and let live" until one day I walked out my
           | door and a wasp flew directly over and stung me without
           | provocation. So I got some insecticide and got rid of the
           | nest.
        
       | praveen9920 wrote:
       | Stepping back a little and try to projecting that logic onto
       | humans, are we more like super organisms? Interestingly, Our
       | social constructs does have similarities of both superorganism
       | and non- superorganism
        
         | xyzzy_plugh wrote:
         | It's even more extreme than that. If you took a random sampling
         | of humans to a version of Earth that had never experienced
         | humans, it would not go well. We're heavily dependant on
         | society and modern technology. We've evolved together.
         | 
         | E.g. smaller birth canals and C-sections.
        
       | cryptonector wrote:
       | Honeybee queens are the only honeybees with stingers that don't
       | die when they sting. That's because the queen bee's stinger has
       | no barbs, and the reason for that is that the queen must not die
       | easily, and she _must_ use her stinger, so if she 's going to
       | survive at all her stinger has to not have barbs. The queen
       | almost certainly has to use her stinger when she exits her
       | cocoon: to kill ther other queens that are about to hatch or have
       | hatched already. She also has to possibly use her stinger when
       | she goes out to mate (though she does go with attendants who will
       | defend her if attacked).
       | 
       | I was surprised not to find mention of this in TFA.
       | 
       | > A honey bee dies when it stings you because its stinger is
       | covered in barbs, causing its abdomen to get ripped out when it
       | tries to fly away. And surviving with your guts spilling out
       | everywhere is pretty bloody hard.
       | 
       | There's another interesting detail here: when the worker tries to
       | fly off after stinging, she has to try _really_ hard because the
       | barbs hold the stinger in place, and trying hard causes two
       | things to happen:                 - noise that attracts other
       | workers         to attack the same creature              -
       | spreading of the dying bee's         distress pheromones that
       | also         attract other workers to sting         the same
       | creature
       | 
       | So when you get stung by a bee near other bees you will be in
       | trouble. That's how you go from one sting to hundreds. And
       | hundreds is enough to kill a human. That's why you don't go near
       | a hive without protection. Being in or near a swarm is safer than
       | being near a hive: the bees in a swarm don't have much (larvae,
       | honey) to protect, so they don't attack.
        
         | abnercoimbre wrote:
         | Useful reminder nature is terrifying even at its smallest. I'm
         | a little surprised this wasn't taught to me in school.
        
           | nejsjsjsbsb wrote:
           | Yeah: caterpillar parasites
        
         | teeray wrote:
         | The queen bee is a formidable final boss with a bad-ass origin
         | story.
        
           | cryptonector wrote:
           | She's also her daughters' slave. They make her work (lay
           | eggs). They decide when to make new queens. They decide when
           | to swarm with the old queen, and when they do they put her on
           | a diet first so she can lose weight so she can fly (they
           | won't let her eat much for two weeks), and they'll push her
           | out of the hive when the time comes.
           | 
           | Humans only really get stung by queen honeybees when
           | manipulating them. Normally the queen will be inside the hive
           | and stay inside the hive except once or twice early in her
           | life when she goes out to mate.
        
             | stavros wrote:
             | Why does she go out to mate? Aren't the drones in the hive?
        
               | geoka9 wrote:
               | Maybe for the same reason we don't marry our cousins? :)
        
               | Jedd wrote:
               | I think two answers, though as per TFA there's probably a
               | lot more behind this:
               | 
               | First, she wants to get the _best_ genetic material, and
               | flying high is one of those tests for drones to pass, and
               | 
               | Second, needs to acquire genetic material from different
               | bloodlines, hence does the business with several drones
               | (obviously not an option within the hive).
        
               | cryptonector wrote:
               | The drones in the hive are her brothers. She needs more
               | gentic diversity, so she goes out to mate with other
               | colonies' drones. Her attendants know where to go find
               | them because they've scouted them before.
        
               | ternnoburn wrote:
               | The drones are also clones (?) of the prior queen. They
               | are unfertilized eggs, so they contain a genetic copy of
               | half the queen's DNA. Mating with a drone from you hive
               | would be like mating directly with your genetic mother.
               | It's going to result in real bad inbreeding real quickly.
               | Especially since queens mate a single time, and live for
               | only a few years.
        
               | cryptonector wrote:
               | They're like half-clones, yeah. Queens can mate more than
               | once though, but usually just once, and if more than once
               | just twice. A well-mated queen will have sperm from a
               | dozen different drones.
        
               | stavros wrote:
               | Ahh, thank you, I thought she mated with her own drones
               | and was wondering how they got any genetic diversity like
               | that.
        
           | inopinatus wrote:
           | it's a Game of Drones
        
             | fuzztester wrote:
             | Do you Lisp?
        
         | ianbicking wrote:
         | This maybe points to another theory (which may be entirely
         | wrong, I'm just guessing!): honeybees die because they aren't
         | supposed to attack each other. Like they can't be aggressively
         | selfish because they'll just die in the process.
        
           | cryptonector wrote:
           | Honeybees do attack other colonies' honeybees. Africanized
           | honeybees definitely do it. As someone else points out the
           | barbs don't get stuck in insects, but do get stuck in mammals
           | (and presumably birds too?).
        
             | fuzztester wrote:
             | Checked them. Found:
             | 
             | https://en.m.wikipedia.org/wiki/Africanized_bee
        
               | cryptonector wrote:
               | > Africanized honey bees are typically much more
               | defensive, react to disturbances faster, and chase people
               | further (400 metres (1,300 ft)) than other varieties of
               | honey bees.
               | 
               | My bees will chase me about 200 yards, and probably more
               | if I had to go further to go inside a building (they
               | don't like dark places). They lay ambushes, too. They'll
               | wait outside the building I go into and will attack again
               | if I go out.
               | 
               | Well, they used to anyways. Since switching to top-bar
               | hive bodies they're much nicer.
        
         | treis wrote:
         | >The queen almost certainly has to use her stinger when she
         | exits her cocoon: to kill ther other queens
         | 
         | I think this is one of the more interesting differences. Plenty
         | of species operate in groups. But its usually a dominant male
         | with a harem. Bees and the like are unusual in that it's a
         | dominant female.
         | 
         | I think it's related to the ease of reproduction. The females
         | put relatively little into their off spring compared to a lion
         | or even birds. It lets them to be essentially autonomous in
         | reproduction which allows them to create offspring that are
         | more like limbs.
        
           | ajuc wrote:
           | There's 13 000 species of ants and 6000 species of mammals,
           | not to count all the other insects.
           | 
           | We are the unusual ones.
           | 
           | BTW there are mammals that live in insect-like groups - for
           | example https://en.wikipedia.org/wiki/Naked_mole-rat#Roles
        
           | cryptonector wrote:
           | There are bees that are monogamous, where the female and the
           | male act a lot more like birds as far as mating and young
           | rearing goes, and there are eusocial bees that where the
           | female absolutely dominates.
           | 
           | Honeybees keep very few males (drones) around, and in the
           | Fall they push them out of the hive and let them die of
           | exposure to the elements. Honeybee workers work themselves to
           | death. The queen is their slave. It's a pretty crappy life,
           | but they make wonderful honey, and they collect wonderful
           | propolis.
        
             | zamfi wrote:
             | > It's a pretty crappy life
             | 
             | By human standards, sure.
             | 
             | Weird take I know, but since only one female bee in the
             | hive passes along her genes (which are shared with the
             | other bees), it's a very different incentive structure.
        
             | ChrisMarshallNY wrote:
             | From what I can tell, the entire hive is "the" organism,
             | and individual bees are like cells in the body. There was
             | an episode of _Cosmos_ , where Neil deGrasse Tyson
             | described bees as the other of two major intelligences on
             | Earth, and suggested their structure as a model for alien
             | visitors[0].
             | 
             | If we judged ourselves by the cells in our body, we'd
             | probably conclude that humans aren't "happy."
             | 
             | White blood cells basically run suicide missions against
             | intruders, so they are sort of like swarming worker bees.
             | 
             | [0] https://m.youtube.com/watch?v=ubE9hjrsHmI
        
         | ivankelly wrote:
         | The bees in a swarm have filled themselves with food before
         | swarming, so they're stuffed, so it's hard to flex the abdomen
         | to make a sting
        
           | cryptonector wrote:
           | Yes, there is this too.
        
         | the_af wrote:
         | Very interesting!
         | 
         | Don't wasps have a similar "swarm attack" mode that doesn't
         | require individual wasps dying to spread pheromones? Something
         | _in_ the sting /venom itself?
         | 
         | I've been stung by wasp swarms twice, in the same area (they
         | were protecting their nest, and please don't ask why it
         | happened twice... we humans do indeed stumble with the same
         | stone twice!). The wasps were very aggressive stinging near the
         | same location in my arm, and it hurt a lot. I was stung in the
         | same body part by the swarm, not in random locations.
         | 
         | No wasps were harmed during this accident.
        
           | cryptonector wrote:
           | Wasps absolutely have attack pheromones, and they absolutely
           | spread those when they get injured (in the same way that bees
           | do when they're injured) and when they attack (but here they
           | might be able to do it "consciously"?).
           | 
           | What's nice about wasp stings is that because the stinger
           | with the venom pouch and pump doesn't get stuck in your skin
           | you get a much smaller dose than with a bee sting.
           | 
           | When I've been stung by bees where the stinger didn't get
           | stuck in my skin the sting was no big deal. When the stinger
           | gets stuck in my skin it's much worse.
        
           | fuzztester wrote:
           | Wasp bites are a lot more painful than mosquito bites.
        
             | the_af wrote:
             | Mosquito bites are not painful at all to me.
             | 
             | I've been stung by a bee only once, and it wasn't
             | particularly painful (but not nothing, of course).
             | 
             | As I mentioned above, I've been stung by swarms of wasps
             | twice and it was _very_ painful.
             | 
             | I hear yellowjacket stings are extremely painful, but even
             | though I've been near them I've never been stung...
             | fortunately.
        
               | fuzztester wrote:
               | >Mosquito bites are not painful at all to me.
               | 
               | interesting. I had a friend who was like that. once he
               | invited me to a jazz concert near a river in the city
               | where we both lived. river = mosquitoes. it was around 7
               | to 8 p.m. there was a shitload of mosquitoes around, and
               | they were biting me and other people like mad, but he
               | seemed completely unaffected. I asked him, aren't you
               | being troubled by them? he said no.
               | 
               | I had had enough, and ran away from both the concert and
               | those mosquitoes.
               | 
               | I have been stung by wasps a few times, and it is a lot
               | more painful than a mosquito bite.
               | 
               | yellowjacket seems to be an American term. I had to look
               | it up.
               | 
               | https://en.m.wikipedia.org/wiki/Yellowjacket
        
               | the_af wrote:
               | Don't get me wrong, I hate mosquitoes and don't like
               | being stung (both because of dengue fever risks in the
               | case of aedes, and because it itches a lot afterwards),
               | but I never find their bite painful. I suppose it depends
               | on the kind of mosquito? I do know -- cause I've been
               | bitten -- that _horseflies_ hurt like hell, because their
               | mouth parts have evolved to pierce cattle skin!
               | 
               | "Yellowjacket" is the translation I found for a kind of
               | wasp we have in Argentina's Patagonia, where we call them
               | "chaquetas" (Spanish for jackets). I think they might be
               | imports and not native though, but now they are _very_
               | frequent in our region.
        
           | ChrisMarshallNY wrote:
           | Depends on the wasp/bee species. Not all of them swarm. Also,
           | ants are related to wasps, and you will see similar behavior
           | with them.
           | 
           | I had a friend, when I was a kid, that shot a white-faced
           | hornet nest (not actually hornets -they are big yellowjackets
           | -even worse) with a BB gun, from, like, 50 meters away.
           | 
           | The hornets figured out who shot their nest, and swarmed him.
           | May have just been that they attacked any nearby critters,
           | but he certainly paid for his folly.
           | 
           | White-faced hornets are better to have in your yard, than
           | yellowjackets. They are a lot less aggressive, only attacking
           | if their nest is at risk. They also eat yellowjackets, so you
           | have hornets _or_ yellowjackets, but not both.
        
             | the_af wrote:
             | Wow. Those hornets seem surprisingly clever.
             | 
             | I know Yellowjackets, we have them in Argentina. Their bite
             | is said to be _extremely painful_ but I haven 't been
             | bitten yet despite some pretty reckless behavior on my
             | part.
             | 
             | One nice afternoon I was having a snack in a cafeteria, by
             | the open window, and a bunch of yellowjackets started
             | dipping in my drinks and my pie. Apparently they like raw
             | meat (locals use it to drive them away from their own food)
             | but I only had sweet stuff on my table. I got annoyed and
             | killed a bunch of them with my bare hands, with no
             | repercussions (killing dangerous insects with your fists or
             | palms is surprisingly easy if you strike fast against a
             | flat surface -- obviously I waited till they moved away
             | from my stuff).
             | 
             | I was later told this was a terrible idea because their
             | bite is very painful (see above: I'm not the smartest about
             | bugs and have been bitten more than once due to
             | recklessness!).
        
               | ChrisMarshallNY wrote:
               | White-faced hornets[0] are a lot more painful than
               | yellowjackets[1] (they are about twice as big, but pretty
               | much the same configuration).
               | 
               | I don't think yellowjackets will swarm, if a forager gets
               | killed. Don't step on one of their nests, though (they
               | dig holes). Also have a friend who did that, once. Also,
               | if you get close to one of their nests (hard to see),
               | they might swarm you.
               | 
               | The biggest difference, is that yellowjackets are much
               | more aggressive foragers, so it's easy to piss off
               | individuals. I had [yet another] friend that imbibed one
               | that went into his can of soda. He tells me the
               | experience left his lip swollen like a cartoon, for
               | several days.
               | 
               | [0] https://en.wikipedia.org/wiki/Dolichovespula_maculata
               | 
               | [1] https://en.wikipedia.org/wiki/Yellowjacket
        
               | cryptonector wrote:
               | Neither bees nor wasps will swarm if far from their nests
               | mainly because there won't be many as they forage in a
               | solitary manner.
               | 
               | But when hornets attack as a group, then they swarm, and
               | they use their pheromones to aid in swarming. See:
               | 
               | https://www.youtube.com/watch?v=onq9ixC7OEg
               | https://www.youtube.com/watch?v=D00mEROqKwU
               | https://www.youtube.com/watch?v=vx2EA_ooCf0
               | 
               | where beekeepers use sticky sheets to get one hornet and
               | then all the other hornets will join the one because they
               | are all attracted to its aid by its pheromones.
        
         | the_af wrote:
         | > _so if she 's going to survive at all her stinger has to not
         | have barbs. The queen almost certainly has to use her stinger
         | when she exits her cocoon: to kill ther other queens that are
         | about to hatch or have hatched already_
         | 
         | But in this particular case the queen is no different from
         | worker bees, right? They wouldn't die either from stinging
         | other bees...
        
           | cryptonector wrote:
           | My understanding is that the barbs in the stingers don't get
           | stuck in insects, but I don't know if that's a fact. At any
           | rate the queen is sometimes irreplaceable, so she needs every
           | survival advantage possible.
        
         | crazydoggers wrote:
         | Great point about the queen bee. The queen also uses her
         | stinger inside her own hive. If the hive believes she is
         | nearing the end of her reproductive abilities, they'll rear
         | other queens. She has to defend her position and quickly goes
         | to kill the new queens when they emerge or she'll be replaced.
         | Beekeepers generally replace their queens every 2 years or so,
         | and have to remove the old queen to prevent this behavior as
         | well.
         | 
         | > That's why you don't go near a hive without protection.
         | 
         | This is true and recommended for just about everyone.. however,
         | I have known some beekeepers that can do it. If you smoke the
         | bees well, they'll get calm enough that they won't sting. It's
         | thought to be likely because the smoke covers pheromones, and
         | also because the smoke causes the bees to begin hoarding honey
         | in case they need to quickly leave a burning hive.
         | 
         | I was never brave enough to attempt it, however during my time
         | beekeeping I was amazed at how magical smoke was. Picking up
         | handfuls of bees in their hive and not being swarmed feels like
         | a super power.
        
           | cryptonector wrote:
           | > I was never brave enough to attempt it, however during my
           | time beekeeping I was amazed at how magical smoke was.
           | Picking up handfuls of bees in their hive and not being
           | swarmed feels like a super power.
           | 
           | I'm not brave enough to attempt it either, except -maybe,
           | someday- with a) smoke, b) a veil instead of a suit, c) nukes
           | rather than mature hives, d) in top-bar hive bodies rather
           | than Langstroth boxes. Fuck Langstroth boxes -- you can't
           | help but kill bees in those boxes, and the moment you kill
           | one you're in trouble. Top-bar hives make it much much easier
           | to not kill any bees.
        
         | cryptonector wrote:
         | BTW, when I get stung, if the stinger doesn't get stuck I
         | sometimes don't even notice till much later -- this happens
         | when I get stung through my gloves. If the stinger does get
         | stuck in my skin the first thing I notice is the buzzing of the
         | bee that stung me. The frequency of the buzzing of the bee that
         | stung me and that is trying to rip off the stinger is
         | absolutely terrifying because I know what comes next: a dozen
         | more bees will flock right to where the one bee stung me and
         | will all try to sting me, and since the one bee was able to it
         | might be the case that her sisters will succeed as well. That
         | buzzing spreads her injury pheromones, and the frequency of its
         | sound also acts as a very loud and clear signal to her sisters.
         | The buzzing lasts about 1 second or less; the pain from the
         | sting comes a second or two later. It takes her sisters about 1
         | to 2 seconds to find the bee that stings successfully. Getting
         | stung by a dozen bees at once is panic inducing, and the
         | swelling that will create will take two weeks to subside. I was
         | stung 13 times at once one time, and 5 times another time. It's
         | no fun.
        
           | jimmaswell wrote:
           | Why are bee stings such a common occurance to you? Are you a
           | beekeeper?
        
             | cryptonector wrote:
             | I am.
        
           | majgr wrote:
           | What I noticed is that pitch of buzzing is higher when bee is
           | attacking, not after sting. Higher pitch means probably
           | higher frequency of wings, so bee has more velocity and it is
           | more nimble. When I was kid visiting frequently a place with
           | 100 hives in a summer. I started to recognize that pitch of
           | attacking bee and learned to keep head lower, between arms.
           | Sometimes I was chased by a bee several tens of meters. It is
           | good to use cover of some trees or bushes. I believe bees are
           | aroused by smell of venom itself. Sometimes, when hive is
           | opened, bees turn their abdomen higher, towards opening
           | showing stings. I think I saw that drops of venom forms
           | around end of sting. Smoke particles, probably bonds to venom
           | vapors, neutralizing its influence on other bees, because
           | after couple smoke puffs venom smell is not noticeable any
           | more. Also, smoke causes bees to change their process, from
           | ,,intruder alert", to ,,tree is burning", and are turning on
           | ventilation. Each summer, first sting swelled the worst, but
           | consecutive ones were less so, so I think some tolerance
           | develops, but it won't last a year. The worst is being stung
           | in fingertip, because it causes nail deformation, for some
           | time.
        
             | cryptonector wrote:
             | Oh for sure, they do buzz differently when they are in
             | protec/attac mode.
        
             | cryptonector wrote:
             | It's not just ventilation. When they smell smoke they also
             | go eat a bunch of honey so they can swarm away with the
             | queen and start over elsewhere.
        
         | WaitWaitWha wrote:
         | Nice, another beek!
         | 
         | I have had a hive where I had queens that lived quite fine
         | together. Weirdest hive as one of them was a _caucasia_ and the
         | other a _cordovan_. Never did get stung with those ladies.
         | 
         | Mine was by accident, but I hear that some beeks do it since
         | cannot get the genetic combination, they just do it as a
         | mechanical combo, to get preferred traits. Like _buckfast_ and
         | _russian_ to help with swarming and defensiveness.
        
           | cryptonector wrote:
           | I've heard of queens co-existing, yeah, but that must be very
           | rare. I wonder if somehow they either smell close enough to
           | the same that they and the workers don't notice, or if they
           | are sufficiently different that they don't recognize their
           | smells as those of other queens.
        
         | credit_guy wrote:
         | For some reason I was never afraid of bees. I have a cousin who
         | keeps bees, and she showed me a bee hive once. Neither of use
         | used any masks or gloves. I remember when she showed me the
         | queen, and I was pointing at it to make sure I got it right,
         | she told me to not touch her, because then the other bees will
         | sense the foreign smell on her and will kill her. In any case,
         | I was surrounded by thousands of bees, but none stang me or my
         | cousin. There's some urban legend that bees smell you, and
         | depending on the smell they decide to attack you or not. I
         | suppose I drew the lucky lottery ticket.
        
         | ciupicri wrote:
         | Does TFA stand for "the f...ine article"?
        
           | cryptonector wrote:
           | And/or the fine author.
        
             | permo-w wrote:
             | the featured article
        
       | cruftbox wrote:
       | Hobby beekeeper here.
       | 
       | Worker bees dies when they sting a person, because the stinger
       | and venom pump remain when they fly off, ripping their abdomen
       | open.
       | 
       | The purpose of this is that the venom pump continues to function
       | after they have left, making the sting as painful as possible.
       | 
       | Honeybees are a superorganism, where the survival of the colony
       | supersedes the survival of any individual bee.
        
         | isityouyesitsme wrote:
         | your comment was excellent.
         | 
         | I couldn't read past the article's pretentious opening.
        
         | majgr wrote:
         | Sometimes I observed how it worked after being stung by a bee.
         | It is better to remove sting using force from one side. When
         | two fingertips are used to remove a sting all venom is being
         | pushed into the wound.
        
       | _orz_ wrote:
       | The title is a bit misleading as it really doesn't explain why
       | bees die when they sting in the sense of a causality. The article
       | itself mentions that the stinging mechanism bees use, is itself
       | not a prerequisite for how they are organized as wasp use a
       | different one. Very interesting read though.
        
       | gunian wrote:
       | Kind of got me thinking is there any tool or is someone working
       | on something that can parse a DNA and give you the end result for
       | any species? If not what's the main challenge
       | 
       | Would be a cool challenge for all the quantum supremacy folk
        
       | supernova87a wrote:
       | The interesting question to me is, "does the _bee_ know it's
       | going to die if it stings you?"
       | 
       | And therefore, acts judiciously in deciding when, if ever, to
       | sting? So that it only does it when it's life-threateningly mad
       | at something?
        
       | buggy6257 wrote:
       | Because Melissa pissed off Zeus.[1]
       | 
       | [1] https://crawliomics.wordpress.com/2019/06/12/zeus-the-
       | honeyb...
        
       | lisper wrote:
       | > The haplodiploidy hypothesis, though nice, is not without its
       | problems. Firstly, there are plenty of species with haplodiploid
       | genetics which do not form eusocial colonies.
       | 
       | That is not a problem. Reproductive fitness must always be
       | measured relative to an environment. Just because colony life is
       | a successful strategy for haplodiploid species in some
       | environments does not mean it will be a successful strategy in
       | all environments. You can see this in other species. Female lions
       | live mostly in cooperative groups while male lions and all other
       | large cats (leopards, cheetahs, and tigers) are solitary.
        
       | YeGoblynQueenne wrote:
       | To prove Mr Feynman right: because I stomp on it.
        
       | JohnHaugeland wrote:
       | For the record, almost no bees die when they sting you. This is
       | just a weird myth that people repeat nonstop.
       | 
       | This _only_ happens to honeybees, and _only_ when they 're
       | stinging humans. Most skin is much thicker than ours, more
       | similar to our feet callouses. The stinger isn't supposed to go
       | in. That's closer to a sword's foil.
       | 
       | The reason they panic when it happens is it's not supposed to
       | happen.
       | 
       | The stingers are mostly meant for combat with other insects. You
       | can watch these fights on youtube. They never, ever end with the
       | bee self eviscerating.
       | 
       | This is a simple example of people trying to look smart by
       | explaining things which, when you check, just aren't true.
        
       | ErigmolCt wrote:
       | Isn't that just so unbearably sad? These little creatures work
       | tirelessly their whole lives, buzzing from flower to flower,
       | pollinating the plants that sustain ecosystems and our food
       | supply. They barely get to live a few weeks as worker bees, and
       | then if they ever need to sting to protect their hive... that's
       | it. Their life ends in an act of ultimate devotion.
        
       | leobg wrote:
       | They can actually survive if you let them.
       | 
       | Here's a video of a guy filming himself being stung by a bee,
       | then watching as the bee spins around, works its stinger loose,
       | and flies away:
       | 
       | https://youtu.be/G-C77ujnLZo
        
         | mirzap wrote:
         | Bees don't die right away; they all fly off after stinging, but
         | they will eventually die for sure.
        
       | apopapo wrote:
       | This entire article is flawed in my opinion. Bees are not
       | "suicidal". They are not meant to die after they sting. It's just
       | that they have not adapted to stinging mammals with elastic skin
       | (and probably will not). Bees can absolutely live after stinging
       | if they manage to dislodge the stinger from the skin.
        
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