[HN Gopher] Mitochondria and the origin of eukaryotes
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       Mitochondria and the origin of eukaryotes
        
       Author : Tomte
       Score  : 142 points
       Date   : 2022-06-18 11:51 UTC (11 hours ago)
        
 (HTM) web link (knowablemagazine.org)
 (TXT) w3m dump (knowablemagazine.org)
        
       | Koshkin wrote:
       | If you are curious about these things, do yourself a favor and
       | dive into Alberts et al. _Molecular Biology of the Cell_. I can
       | 't recommend it enough.
        
         | feet wrote:
         | Very solid text book +1
        
         | wrycoder wrote:
         | Alberts et al. Essential Cell Biology is a bit less advanced,
         | but I'm finding it very complete, just not quite so exhaustive
         | (pun intended). It's still 700 pages, so you'll get your
         | money's worth (if you buy a used copy).
        
       | [deleted]
        
       | sonicggg wrote:
       | Evolution did a terrible job there. So many diseases linked to
       | mitochondrial dysfunction nowadays. We have this amazingly
       | complex machinery, and a single point of failure, no redundancy
       | whatsoever.
        
       | lock-the-spock wrote:
       | An accessible and incredibly insightful read 100% on this topic
       | is "Power, Sex, Suicide: Mitochondria and the Meaning of Life"-
       | despite the flashy title a thorough scientific read and analysis,
       | working through various possible arguments and teaching
       | fundamentals along the way.
       | 
       | My favourite book I read in the past five years or so
       | 
       | https://en.wikipedia.org/wiki/Power,_Sex,_Suicide
        
         | aesch wrote:
         | I haven't read that one but after reading The Vital Question I
         | will recommend anything with Nick Lane's name on it. There are
         | very few people who have thought about the origin of life as
         | deeply as he has.
        
           | code_biologist wrote:
           | Read it as soon as you have the time. It is every bit the
           | equal of The Vital Question.
        
         | soramimo wrote:
         | Thanks for sharing, just ordered.
        
       | alangibson wrote:
       | > For billions of years after the origin of life, the only living
       | things on Earth were tiny, primitive cells resembling today's
       | bacteria
       | 
       | I've often used this "fact" as an argument against complex
       | extraterrestrial life. Glad to see it might be wrong after all.
        
         | griffzhowl wrote:
         | Nothing in the article contradicts that 'fact'. The question is
         | only whether the development of eukaryotes involved other
         | significant steps in addition to the mitochondrial
         | endosymbiotic event, or whether the latter event was all that
         | really mattered. In both cases, subsequent development of
         | eukaryotes and hence multicellularity depended on the seemingly
         | very rare endosymbiotic event.
        
         | majkinetor wrote:
         | This is Nick Lane's argument - that complex cell happend only
         | once on Earth (no convergent evolution) via this symbiotic
         | accident and hence it's highly improbable if not almost
         | impossible.
         | 
         | As usual, people always seem to foreget that in this universe,
         | what happened once, happens all the time and we are not
         | special.
        
           | prox wrote:
           | And that is if there is only one universe, which might not be
           | so special as well.
        
           | ssl232 wrote:
           | Nick Lane's book The Vital Question is a masterpiece. I
           | reckon he's figured out the origin of life there. I hope he
           | gets on some podcast like Lex Fridman to talk about it all,
           | because his ideas are just waiting to explode into the public
           | imagination.
        
             | aesch wrote:
             | He was just on Sean Carroll's podcast Mindscape, good
             | listen: https://www.youtube.com/watch?v=9IZ_CVu2M68
        
           | marcosdumay wrote:
           | > that complex cell happend only once on Earth
           | 
           | Well, it's quite possible that once it happens, it voids any
           | chance of it happening again just because life filling that
           | niche already exists. If so, it only happens once, and this
           | says absolutely nothing about the odds of that first
           | occurrence.
           | 
           | > what happened once, happens all the time
           | 
           | That's a fallacy people do all the time here when talking
           | about exobiology. It's basically states that all small
           | numbers and all big numbers are alike, and that all small
           | numbers are like the inverse of all big numbers.
        
             | zeroonetwothree wrote:
             | Most other features evolved independently several (or even
             | many) times so this is not a good argument. It's not as if
             | once there was a eukaryote it instantly spread across the
             | whole planet, taking up every ecological niche. Even today
             | eukaryotes don't exist in certain environments.
        
           | alangibson wrote:
           | We're all just speculating given that were stuck at sample
           | size = 1 until we develop near light speed travel.
           | 
           | I get the convergent evolution point, but so far the story is
           | that 1) life arose very quickly 2) then proceeded to do very
           | little for billions of years. If that's true, then it's bad
           | news for the possibility of complex life being common.
        
             | majkinetor wrote:
             | It only means that complex life is less frequent then
             | single cell life, not that it is uncommon.
        
             | Qem wrote:
             | > 1) life arose very quickly 2) then proceeded to do very
             | little for billions of years.
             | 
             | This is understating the feats of unicelular life. Those
             | tiny organisms essentially terraformed the whole planet in
             | those billions of years, oxidizing nearly all the iron
             | close to earth surface, burying most carbon and then
             | pumping so much oxygen in the atmosphere that at a point it
             | comprised 30% of Earth gaseous envelope by weight. I
             | wouldn't call that very little. To come close we'll need at
             | least to terraform Mars or Venus, to match the feats of our
             | microbial forebears.
        
         | tomrod wrote:
         | Single-celled life is complex. It has the same length of
         | evolutionary time as multicellular life.
         | 
         | We know very little about modes of sentience with N=1. Perhaps
         | life is generally mutualistic.
        
         | photochemsyn wrote:
         | The rise of free atmospheric oxygen and oceanic dissolved
         | oxygen levels were almost certainly a requirement for the
         | development of complex multicellular life. Using oxygen as the
         | electron acceptor in respiration means far more energy can be
         | extracted from reduced fuel molecules (sugars, fats, amino
         | acids, methane, hydrocarbons, ammonia, etc.) than by using
         | species like oxidized sulfur and iron, CO2, or nitrate (NO3)
         | for that role.
         | 
         | This is really where the mitochondria come into play, as they
         | allowed their host to utilize oxygen for this purpose. This can
         | be seen by looking at modern eukaryotes that have reverted back
         | to an anaerobic lifestyle:
         | 
         | > "In lineages of eukaryotes adapted to low oxygen conditions,
         | mitochondria have been drastically reduced, functionally
         | altered and, in one case, completely lost."
         | 
         | "The Origin and Diversification of Mitochondria (2017)"
         | 
         | https://www.cell.com/current-biology/pdf/S0960-9822(17)31179...
         | 
         | Hence, looking for the signature of free oxygen in the
         | atmosphere of exoplanets orbiting distant stars is considered
         | to be a fairly good indicator of the possibility of complex
         | multicellular life of some sort, and at least of an active
         | photosynthetic microbial ecosystem.
         | 
         | (Incidentally, the historical divisions withing academic
         | university departments led to evolutionary biology generally
         | ignoring the importance of early Earth's geochemistry in the
         | evolution of life, as they saw evolution as a kind of
         | cellular/organismal process divorced from the physical
         | surroundings - the latter being the province of the geology
         | department. The renewed interest in exobiology and origin-of-
         | life research has tended to bridge this gap.)
        
           | alangibson wrote:
           | The way I read that is that developing complex life is even
           | more contingent than 'just wait long enough'. Given what we
           | know for sure, you need rather narrow concentrations of
           | specific gasses at least.
        
         | simonh wrote:
         | It's not wrong, at least I don't think this article is arguing
         | that multi cellular life evolved any earlier than we thought.
         | Just that the mechanisms were different than we expected.
        
       | pmags wrote:
       | For those interested but new to role of endosymbiosis in
       | evolution, it's useful to keep in mind that there's strong
       | evidence of an endosymbiotic origin of chloroplasts, another
       | major event in the history of life. Here's a recent review
       | article that highlights some of the key evidence and outstanding
       | questions:
       | 
       | Sibbald SJ, Archibald JM. Genomic Insights into Plastid
       | Evolution. Genome Biol Evol. 2020 Jul 1;12(7):978-990. doi:
       | 10.1093/gbe/evaa096. PMID: 32402068; PMCID: PMC7348690.
        
       | csr86 wrote:
       | How the children inherited the swallowed organism?
        
         | majkinetor wrote:
         | Bacterial division would get some mitos on either side of the
         | dividing cell. Mitos then proliferate on its own.
        
         | frostburg wrote:
         | Presumably like we do - (I'm heavily simplifying the complex
         | regulation involved) the mitochondria have their own DNA which
         | gets duplicated when the cell prepares to undergo mitosis, then
         | the two new cells split the results.
        
           | anton96 wrote:
           | Does that implies that mitochondria also had to evolve at
           | some point to detect duplication of their host cell so they
           | duplicate at the same time?
        
             | reubenswartz wrote:
             | There are lots of mitochondria in a cell (almost always,
             | ignore red blood cells, for example), and the mitochondria
             | have their own mechanisms for detecting when they need to
             | divide (exercise and the resulting stress being one of
             | them, which is why working out makes you more fit).
             | 
             | Also note that the DNA for most of the mitochondrial
             | proteins is in the host cell's nucleus-- the mitochondrial
             | DNA is stripped down to coding for just a few essential
             | proteins.
        
       | Linda703 wrote:
        
       | anuvrat1 wrote:
       | How simple are Prokaryotic Cells really? Like "simple" enough
       | that we understand all the intricate workings of it, or "simple"
       | enough to be made from scratch?
        
         | david_l_lin wrote:
         | Honestly, neither. Biology is complex and messy, and often not
         | at all intuitive.
        
         | Daniel_sk wrote:
         | Imagine giving a scientist in 1900 a modern CPU. He may be able
         | to disassemble and study it, but it's impossible to make it
         | from scratch because it takes a chain of steps that lead to
         | complicated modern factories that then produce the chip. It's
         | hard to skip those steps. We can swap DNA in living cells and
         | produce modified cells, but we lack the tools to create such
         | complicated and microscopic structures in a pure synthetic way.
        
       | wheelerof4te wrote:
       | A neat fairy-tale but, with all due respect, scientist don't know
       | anything that happened a billion+ years ago.
       | 
       | This is a little more than a tabloid article.
        
       | ncmncm wrote:
       | If they find cells lacking some organelle, it will be hard to say
       | whether it was never acquired, or was later abandoned.
       | 
       | One clue would be leftover genes from when it was there.
       | Eukaryotes often find it hard to prune out excess DNA, vs. just
       | turning it off.
        
         | edgyquant wrote:
         | DNA doesn't help as much as you think with single celled
         | organisms thanks to HGT. Bacteria, at least, share DNA and thus
         | we can't use it to determine a linear evolutionary sequence.
        
           | ncmncm wrote:
           | Eukaryotes don't do so much of that.
        
           | jjtheblunt wrote:
           | HGT?
        
             | [deleted]
        
             | reubenswartz wrote:
             | Horizontal gene transfer.
             | 
             | Because biology isn't complicated enough. :)
        
         | 323 wrote:
         | > _If they find cells lacking some organelle_
         | 
         | A most famous example is red blood cells, which are emptied of
         | most organelles to make space for hemoglobin.
        
       | [deleted]
        
       | CRUDite wrote:
       | If we assume single celled life is common, and exists in either
       | all star systems, or in all systems without a hot Jupiter close
       | to the star; then given the length of time involved here before
       | eukaryotes, can we estimate how many planets in the galaxy have
       | eukaryotes? Or how long it will be before some do? Presumably
       | increasing the length of time increases the probability of
       | occurrence as would increasing the number of stars involved.
       | Though how can we know if the event was statistically likely
       | after that amount of time or whether we are deviations from the
       | centre of the distribution (other than observing the quiet
       | galaxy). Surely we can have a rough idea now of where we stand
        
         | JumpCrisscross wrote:
         | > _can we estimate how many planets in the galaxy have
         | eukaryotes?_
         | 
         | Not yet. In the history of life on Earth, this has happened
         | once. Knowing what we know about cellular biology, it's
         | stupidly unlikely. Beyond our present theories' ability to
         | quantify.
         | 
         | By the way, I think this is one of--if not the--great filters.
         | It's unlikely to happen, to not promptly get smote by its
         | primordial planet's tantrums and to get it so right it
         | perpetuates for billions of years.
        
           | echelon wrote:
           | > Not yet. In the history of life on Earth, this has happened
           | once.
           | 
           | 1) That we know about.
           | 
           | 2) Not unlike startups vs. established business, any newly
           | emerging "eukaryotes" have to out-compete the already-evolved
           | incumbents, which are already quite good at harnessing
           | energy. You're much more likely to find success in business
           | than in an entirely new evolutionary branch, though I doubt
           | biological "gray goo" is outright impossible [1].
           | 
           | [1] Reverse chirality autotrophs sound like a scary sci-fi
           | novel plot https://news.ycombinator.com/item?id=28038505
        
             | ajuc wrote:
             | Additionally the environment changed significantly since
             | then (for one example oxygen which was highly toxic to most
             | forms of life that existed back then is now over 20% of
             | atmosphere).
             | 
             | > Reverse chirality autotrophs sound like a scary sci-fi
             | novel plot
             | 
             | Very ice-9-like.
        
             | yyyk wrote:
             | >Reverse chirality autotrophs sound like a scary sci-fi
             | novel plot
             | 
             | I doubt this. 'Not being digestible' is very far from
             | 'being invulnerable' or even 'being able to spread
             | quickly'. The kingdom of life has many ways to kill stuff,
             | ways which don't care about chirality, and our typical
             | R-sided lifeforms have all the evolutionary 'motivation' to
             | come up with new ways just the off the competition. That's
             | before humans get into the picture, which we have the tech
             | to do.
             | 
             | There may be an accumulation of non-digestible stuff until
             | nature reaches a balance. However, there's a very large
             | recent accumulation of non-digestible materials called
             | 'plastics', and while somewhat harmful, they're not a life-
             | ending threat. Nature is already finding ways to process
             | these materials[0].
             | 
             | [0] https://en.wikipedia.org/wiki/Plastic_degradation_by_ma
             | rine_...
        
               | echelon wrote:
               | Sure, but I did say "sci-fi". And I think there are a lot
               | of unaddressed points.
               | 
               | Plastics don't self-manufacture. You might not be able to
               | control the rate.
               | 
               | Just because you kill something doesn't mean you break
               | down its carbohydrates. Reverse chiral organism skeletons
               | could bioaccumulate and we could have a situation similar
               | to the Carboniferous.
               | 
               | Someone might be able to synthesize a bacteria in the lab
               | given enough time and effort from an organism that
               | proliferates quickly. It doesn't have to capture all the
               | carbon. Just out-compete a keystone species. Plankton,
               | mycorrhizae, etc. Or attack a large percentage of the
               | plant biomass.
        
               | yyyk wrote:
               | >Plastics don't self-manufacture. You might not be able
               | to control the rate.
               | 
               | The rate is limited by the process. Since no precursors
               | exist, it must 'self-manufacture' from scratch. This has
               | inherent limits even before introducing competition for
               | food, poison, predators that eat you even despite them
               | not being able to really digest, etc.
               | 
               | >Just because you kill something doesn't mean you break
               | down its carbohydrates. Reverse chiral organism skeletons
               | could bioaccumulate and we could have a situation similar
               | to the Carboniferous.
               | 
               | So you don't break it down. Nature will have plenty of
               | time to adapt. Humans will step in if needed.
               | 
               | >Someone might be able to synthesize a bacteria in the
               | lab given enough time and effort from an organism that
               | proliferates quickly.
               | 
               | That's an incredibly messy way - create an entire
               | L-chiral biochemistery - to get a weapon which doesn't
               | have a setting between 'kill everything' and 'do rather
               | little' (IMHO, the second being much likelier). There are
               | far worse and more directed things one can do with a lab.
               | Even the absurd 'kill everything' goal is far more likely
               | to be reached in different ways.
        
           | sterlind wrote:
           | per the article, endosymbiosis has happened a bunch of times.
           | multiple different kinds of chloroplasts, several
           | prokaryotes, a parasite etc.
           | 
           | this was all when eukaryotes engulfed prokaryotes, but still,
           | how does this mean unlikely? it seems imminently likely,
           | since.. it happened a bunch of times.
           | 
           | seems to me like prokaryotes evolve a strategy of engulfing
           | others for their resources, then one day engulf a prokaryote
           | infected by a virus, which transfers DNA across, rinse and
           | repeat.
           | 
           | how is this more of a filter than abiogenesis?
        
             | simonh wrote:
             | The thing that only happened once on Earth and that's a
             | prerequisite to developing complex life forms is not
             | endosymbiosis, it's life going multi-cellular. They are not
             | the same thing.
             | 
             | Endosymbiosis is not identical with going multi-cellular,
             | and it seems that all but perhaps one of the known
             | instances of endosymbiosis didn't play any role in us going
             | multi-cellular anyway. In fact this article makes the case
             | that it may not have been critical at all.
        
               | gus_massa wrote:
               | Multicellularity looks relatively easy. From https://en.w
               | ikipedia.org/wiki/Multicellular_organism#Occurre...
               | 
               | > _Multicellularity has evolved independently at least 25
               | times in eukaryotes, and also in some prokaryotes, like
               | cyanobacteria, myxobacteria, actinomycetes, Magnetoglobus
               | multicellularis or Methanosarcina. However, complex
               | multicellular organisms evolved only in six eukaryotic
               | groups: animals, symbiomycotan fungi, brown algae, red
               | algae, green algae, and land plants. It evolved
               | repeatedly for Chloroplastida (green algae and land
               | plants), once for animals, once for brown algae, three
               | times in the fungi (chytrids, ascomycetes and
               | basidiomycetes) and perhaps several times for slime molds
               | and red algae._
        
               | simonh wrote:
               | It's relatively easy once you have evolved the
               | biochemical infrastructure to support it, but on Earth
               | that took several billions of years to achieve. There's
               | no way around it, no amount of hand waving how easy it is
               | negates the fact it took billion years of evolution to do
               | it.
               | 
               | Also most of those forms of multicellularity are
               | extremely basic, little more than tangles or sheets of
               | cells, even after hundreds of millions of years of
               | further evolution. That's not likely to get to
               | intelligent life.
        
             | JumpCrisscross wrote:
             | > _how is this more of a filter than abiogenesis?_
             | 
             | Common chemistries get us very close to molecular systems
             | subject to evolutionary pressure. (Simplest: RNA world
             | hypothesis.) We are missing links. But the pathway is
             | plausible.
             | 
             | Chloroplasts, as you mention, are a potent counter
             | argument. But once you have surplus cellular energy,
             | additional endosymbiosis has a lower threshold. Based on
             | current research, all life has a similar mitochondria.
             | Different kingdoms didn't nom their own and go. That
             | uniqueness suggests difficulty.
        
             | kylebenzlee wrote:
             | Yes, this (nonsense) discuassion is when you have computer
             | progammers discuss biology. Is strange because they know
             | nothing but all seem to think they must be experts of
             | evrything because they get paid a lot to sit infront of a
             | computer all day.
        
         | marcosdumay wrote:
         | Well, that's not a good assumption to make. Our most capable
         | life detector1 is analyzing the atmosphere of planets, captures
         | stuff similar to our simplest single celled life. So, it's not
         | a good guess that it's common.
         | 
         | 1 - Actually our second best. The best one is the fact that
         | nobody colonized Earth before we existed, that is tuned in
         | space-faring life.
        
         | kylebenzlee wrote:
        
         | 323 wrote:
         | You make a big assumption, that there must be only two kinds of
         | cells, "simple" and "complex", like on earth.
         | 
         | That could well be an accidental fact. Maybe on some planets we
         | have a gradient of cell complexity.
         | 
         | Also, we don't really know what even simpler kinds existed on
         | earth but were lost, since bacteria, the "simple" kind, it's
         | obviously too complex to have been the first ever life form.
        
           | andrewflnr wrote:
           | That's not even true on earth. Archaea exist, and eukaryotes
           | span a pretty wide range of complexity themselves.
        
         | lkrubner wrote:
         | Maybe, but there are other narratives that are easy to spin.
         | Our solar system has 3 planets all of which might have had
         | single-celled organisms, at some point: Venus, Earth, and Mars.
         | And for the first 3.7 billion years, the 3 planets might have
         | followed a similar path. And then all 3 planets reach old age
         | and basically die, Venus becoming too hot, while Earth and Mars
         | become mostly dead ice covered snow balls. And maybe that is
         | the normal history of most planets, even planets that develop
         | single-celled life.
         | 
         | In that narrative, the emphasis is on the extraordinary re-
         | birth of Earth, after the end of Snowball Earth. Almost
         | everything that we regard as interesting about Earth happens
         | after this late-in-its history revival. That raises some other
         | questions, such as, why did Snowball Earth end? Why does
         | multicellular life take off then, but not before? What is it
         | that makes the Earth/moon system so unusually dynamic that it
         | hasn't settled down to some dead equilibrium, even after 3.7
         | billion years? What allows Earth to have such an extraordinary
         | additional era?
        
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