Testacella haliotidea.
Vitrina pellucida.—The Vitrinas in several ways afford a connecting link between the slugs and snails, having the same tooth-formation and mantle as the former, while the shell cannot contain the whole body. As the name indicates, the shell is like a bubble of clear greenish glass and very delicate. It is small, and found in damp places, coming out mostly at night. Omnivorous, it is often found feeding on dead worms, and, unlike nearly all our earth molluscs, can be found abroad in winter.
Vitrea (Polita) lucida.—This is the largest of our British Hyaliniæ, which are difficult to distinguish. The body of this species is cobalt blue, the apex of the shell is flat, its colour opaque, and the last whorl more expanded than in others. All belong to the sub-genus Polita, and have polished or glossy shells. All love shade and moisture, and should be sought under stones or wood or in moss. They only come out by day when it is wet, a habit they may have acquired from their being a favourite food of birds, 416 having been found in the crop of one nestling Stockdove; while various flies are very destructive to them. This species prefers animal food, and is more gregarious than others. Not common.
Vitrea (Polita) cellaria.—The next largest species is the most common of all. It is fond of cellars (whence its name), and I found it under the stone lid of a manhole in the drain of S. Peter’s Rectory, Walworth—the only shell left in that part of London. It resembles the previous species, but is smaller and has a broader and deeper suture between the whorls, while the foot-sole is paler than the body.
Vitrea (Polita) Rogersi.—Local. Found in dense woods. It is much like both H. cellaria and H. alliaria, and all three smell of garlic, but the last is much smaller than the others. The tentacles in the first are long, and in the third short; while in Rogersi the upper pair are long and the lower very short. It is also the most glossy of all. If put in a box with other small shells it will clean them by cannibalism.
Vitrea (Polita) alliaria, i.e. smelling of garlic.—Often confused by quite good conchologists with the preceding species, but the body is much darker, and the shell smaller and less white below than either cellaria or helvetica. The always present smell is said to protect it from ants. Common, but local, and often a pest in greenhouses and ferneries.
Vitrea (Polita) nitidula.—Common. Less glossy. Marked expansion of the last whorl as it nears the mouth.
Vitrea (Polita) pura.—Like nitidula but smaller, and edge of mantle white instead of dark. More common in the north. Shell thin and dull white.
Vitrea (Polita) radiatula.—Never abundant. Striations on shell give it a radiated appearance when magnified. Animal nearly black.
Vitrea crystallina.—The smallest of the genus. Shell transparent, pearly white. Umbilicus (i.e. the opening in the centre of the underside showing the whorls) very narrow. Subterranean in habit. Whorls, four; whereas H. pura has five; also more compressed.
Euconulus fulvus.—Distinctively pyramidal in shape. Small. Brown. Common under rotten branches and moss in woods. Hardly hibernates.
Zonitoides nitidus.—Chocolate-brown, with no white round the umbilicus (as has H. nitudula). Larger than, but not unlike, H. radiatula. Gregarious. Chiefly found by water; also in damp hothouses. Amphibious.
Zonitoides excavatus.—Its broad and deep umbilicus is quite distinctive. Mainly British. Dislikes lime, and is most plentiful on the coal measures.
We come now to the Helicidæ family and its genus Helix, in which there are various sub-genera of which the name is given in brackets. The shell in this genus can wholly contain the body; the tentacles are always four; the shell conical, and rarely with a depressed spire. The word “helix” is Greek, and means a coil.
Helix (Gonyodiscus) rotundata.—Very common under stones, moss, etc. Circular, flat, with a large open umbilicus. Horn colour with brown markings.
H. (Pyramidula) rupestris, i.e. inhabiting rocks.—Small. Gregarious. Dark brown. Mainly on exposed dry walls and cliffs.
H. (Punctum) pygmæum.—Very small. Yellowish brown and glossy shell. Mainly on moist dead leaves. Not unlike H. rupestris except as to habitat.
H. (Acanthinula) lamellata.—Small. Horn-colour. Epidermis raised into lamellæ or ridges in the line of growth. Mainly northern. Frequents dead leaves, especially beech and holly.
H. (Acanthinula) aculeata.—More common than the former; which it resembles in habitat. Differs chiefly by the ridges being produced into spines.
H. (Vallonia) pulchella.—Tiny. White. Mouth trumpet-shaped. Umbilicus wide. Under stones and at the roots of grass. Its variety costata (which some make a separate species) is strongly ribbed.
H. (Helicigona) lapicida.—Circular, flattened, dark brown, strong white reflected rim to mouth. Large umbilicus. Marked keel, which distinguishes it from all other British land shells. Chiefly on chalk soils. Often on beech tree trunks.
H. (Gonostoma) obvoluta.—Common abroad, but confined in England to a few spots in Sussex and Hants. Circular, flat above, mouth triangular, with a strong pinkish-white rim with three denticles.
H. (Pomatia) pomatia.—Described earlier. Found in Hants, Sussex, Kent, Surrey, Oxford, Gloucester, and Bedfordshire; but very local. Elsewhere it may well be an escape from captivity, or the remains of an attempt (always unsuccessful) to establish a colony. Box Hill and Caterham are two good localities for Londoners. In Kent it has two centres, Charing and Shoreham with their contiguous parishes, but there is a great gap between them, and it is absent from places on the same chalk ridge which are identical in soil and vegetation.
H. (Cryptomphalus) aspersa.—The sub-generic name means that the umbilicus is hidden in adult shells by a fold of the pillar lip; the specific name means sprinkled (with brown blotches); but it may be a slip of the pen, for aspera, or rough, from the rough shagreening of its surface. Five banded, like so many of the Helicidæ, but usually the second and third band unite. No umbilicus. The variety exalbida (chiefly found in Kent and the West) is straw colour and somewhat transparent. Commonly sold for food on the Continent as well as pomatia, which is cultivated in “snail-farms,” but not native in Germany or Switzerland, and in France chiefly found in the coast departments. Insipid; but as nourishing as calf’s-foot jelly. Fond of gardens (whence its common name), but not of gardeners. As most animals are marvellously gifted with a knowledge of what food to eat and what to avoid, it is curious that aspersa will eat voraciously the leaves of the spindle-tree, though this soon poisons them. It is said also that they share with cows and horses the ignorance that the leaves of the yew should be avoided on pain of sickness or even of death.
H. (Cepæa) nemoralis.—As already stated, this is the most brilliantly and variously coloured and diversely banded of all our English land shells with the exception of its very close connection H. (Tachea) hortensis. It is happily very common, and so the attention of beginners should first be directed to this. Thrushes and mice are its great enemies, the former smashing it on some stone which may be found surrounded by the broken shells. The “mouth” or peristome is normally black, the shell larger and stouter than hortensis, in which the mouth is white. When a white-mouthed nemoralis or dark-mouthed hortensis (both rare) is found, the shape of the internal flinty dart at once distinguishes them. In some places both live together: in most one is found and not the other. Nemoralis is fond of sand-hills by the coast, but is chiefly a hedge-snail, and the edges of main roads are preferred because of the greater variety of food, because the traffic scares away their bird enemies, and because the dust gives them abundance of already prepared material for their shells. When, however, the collector comes to a wayside cottage where fowls are kept he need not waste his time in looking for snails in the neighbouring hedge. The more the chicken industry extends and the more the Bird Protection Acts operates, the worse it is for collectors of snails. The banding is probably protective, as in the case of the tiger and the zebra, and renders the shell less visible.
Helix (Cepæa) hortensis.—Rarely found in gardens in spite of its specific name. A hedge-snail. White forms not uncommon, though almost unknown in nemoralis. Though the weaker form, the coalescence of the five bands into one broad one is more common here than in nemoralis. Also the variety with only one band, and that on the periphery, is very common in nemoralis and rare in hortensis. It is more dependent on shade and moisture than its congener. Smells of garlic when immersed in boiling water to be killed. Hortensis is a more northern, and nemoralis a more southern, shell by origin and distribution. There are 89 possible band variations in any normally five-banded shell, and all have been noted in the case of nemoralis, but in hortensis only 61. They are distinguished, for purposes of record and exchange, by numbers. Thus the type is 12345, the usual one-banded variety 00300, the common coalescence of the second and third band is 1(23)45, and when all bands unite (12345). The unicolourous or bandless variations would be 00000.
H. (Arianta) arbustorum.—Local. Usually found in hedges and by ditches on chalk and limestone. Shell globose, brown or yellow, with a check or willow leaf pattern, and a single dark band on the periphery. Lip strong and white. Animal usually nearly black. Very fond of moisture. Anatomically related to A. lapicida, but no external resemblance.
Helix (Theba) cantiana.—First observed in Kent (where it is especially fine and abundant), whence its specific name, but generally dispersed in South and East England. A dull, creamy white shell with a pink tinge, sometimes becoming partially or wholly reddish.
Helix (Theba) cartusiana (first noticed near a Carthusian monastery). Much resembles cantiana, but is much smaller and more smooth. Chiefly found on the downs of Kent and Sussex. Used to be common on Deal sand-hills—now devastated by golf! The tint in this is brown, in the former red.
H. (Hygromia) rufescens.—A flattish, dark brown shell, abundant in the south of England, and not rare elsewhere. Has a semi-lunar mouth with a white internal rib. In gardens seems to prefer violet beds.
H. (Hygromia) hispida, i.e. hairy.—These hairs are deciduous, and the hairless variety used to be considered a separate species under the name of concinna (i.e. neat), but would now be the variety depilata, or bald. Broad and deep umbilicus. Common, except in Ireland. Usually associates with H. rufescens in moist places.
H. (Hygromia) granulata is also hairy with white silky bristles. Yellowish in colour. Shell thin. Local, but abundant where found. Its umbilicus is very small. It falls from its food plants at the least shake.
H. (Hygromia) revelata.—Scantily haired. Globular thin shell. Pale green. Mainly found in Cornwall and South Devon. In cold or dry weather it buries itself rather deeply.
H. (Hygromia) fusca.—Very thin, glossy, brown shell. Local. Hardy, and even active in frost. Chiefly found on nettles, which many shells like as food, though avoiding the commonly associated horehound.
H. (Euparypha) pisana.—First noticed at Pisa. Somewhat like Helicella virgata, but larger, sub-globular, and solid shell, yellowish-white with dark lines or bands. Aperture or mouth yellowish or rosy. Most common in Portugal and Morocco, and all round the Mediterranean, dry places, especially near the sea. In England chiefly confined to Tenby and other parts of Pembrokeshire; also in the Channel Islands. Varies much in tint and markings. Swarms where found; it loves sun and heat. Seems to lend itself better to colonization than most species.
H. (Helicella) itala.—So named by Linnæus, who probably received it first from Italy. Shell almost circular, flat. Umbilicus very large and open. Common on heaths and downs, especially near the sea.
H. (Candidula) caperata.—(The specific name means wrinkled, like a goat’s horn.) Careless of heat or cold. Distinguished from the young of H. virgata by being more depressed, having a larger umbilicus, regular and strong striation, and round mouth with white internal rib. Found under stones and on grass. Common.
Helicella virgata at rest on thistle, natural size.
H. (Heliomanes) virgata (i.e. striped).—A very variable shell. See the illustration above of some at rest on thistles. Local, but very abundant where found. Whitish shell with dark bands, but a yellowish and a white variety usually is found with the type. The most beautiful variety, radiata, is chiefly found in Romney Marsh, and from Hythe to Rye.
H. (Turricola) terrestris.—A Mediterranean species, well established since 1890, in one spot near Dover. A pyramidal shell, greyish, with one dark band on each whorl.
H. (Cochlicella) barbara (i.e. foreign).—Long, conical, whitish, with one dark band. By the sea-coast. In shape somewhat like a Buliminus.
We come now to the Pupa family and its genus Buliminus and its sub-genus Ena. It is represented by:—
Ena montana.—A local and southern shell, conical, slightly glossy, brown. Lip white and deflected. Commonly found on the holes of smooth-barked trees, and it closely resembles the small knobs on beech trunks.
Ena obscura.—Like the former, but much smaller, and found nearly everywhere in England and Wales. Found in hedgebanks, or on beech trunks. Its specific name is derived from its habit of covering itself with a coating of earth, and so becoming inconspicuous.
The plate on p. 47, gives figures of some of our smaller shells, enlarged in most cases so that their distinguishing marks can be seen. The upright line by the side of each figure gives its actual height. The shells as numbered are Helix rupestris, H. pygmæa, H. pulchella, H. lapicida, H. obvoluta, H. terrestris, H. barbara, Ena montana, Ena obscura, Pupa secale, P. anglica, P. cylindracea, P. muscorum, Vertigo antivertigo, V. moulinsiana, V. pygmæa, V. alpestris, V. substriata, V. pusilla, V. angustior, V. edentula, and V. minutissima. Without a magnifying glass it will be seen that it would be very hard to distinguish some of the minute shells, but this enlargement enables us to see the characteristic denticles in the mouth, and the presence or absence of striations on the shell.
Some of our smaller shells. Actual size indicated by the upright line.
Pupa (Abida) secale is named from the Latin for rye, a grain of which the shell more or less resembles. Conical, brown, mouth horseshoe-shaped with eight white denticles. Our largest Pupa. Local, but abundant where found. Prefers calcareous rocks or woods.
Pupa (Lauria) anglica.—Small, ovate, purplish in colour; mouth like that of secale. Lives in moss, mainly in the north of Britain.
Pupa (Lauria) cylindracea.—Small, cylindrical, paler than the last; thick and reflected white lip with one denticle. Abundant. On stones, in moss, under leaves and bark.
Pupa (Jaminia) muscorum.—Common, especially on sandy soils near the sea. Mouth nearly circular, whereas in the two former species it is horseshoe-shaped. The lip is thin and not reflected.
The genus Vertigo (i.e. twisted, the Latin equivalent of the Greek Helix) contains shells even smaller than the Pupæ, about the size of a pin’s head.
Vertigo (Alæa) antivertigo (i.e. not reversed or sinistral, as are V. pusilla and V. angustior). Semi-transparent, glossy, horn-colour, with denticles (as have all except V. edentula and V. minutissima). Found in nearly all counties in moist places.
Vertigo (Alæa) moulinsiana.—Our largest species, though only 2-1/3 millimetres in height. Mainly in marshy places. Not common.
Vertigo (Alæa) alpestris.—Rare and local, chiefly northern. Nearly transparent shell.
Vertigo (Alæa) pygmæa.—Common, and often in colonies at roots of grass and under stones and logs. Not confined to moist places.
Vertigo (Alæa) substriata.—Local. Strongly striated.
Vertigo (Vertilla) pusilla.—Sinistral, as is also
Vertigo (Vertilla) angustior.—Both species rare and local. The former is the larger and broader. In the former the last whorl is broadest, in the latter the penultimate. In the former the mouth is semi-oval, in the latter triangular. In the former the outer lip is very slightly, in the latter very deeply contracted. The former has 6 to 7 teeth, the latter 4 to 5.
Vertigo (Sphyradium) edentula is dextral and without denticles. Perhaps the most common Vertigo. Partial to bracken.
Vertigo (Isthmia) minutissima.—Dextral and without denticles. Smaller, narrower, and more strongly striated than edentula, but rarer. All the Pupæ should be examined with a magnifier.
Balea perversa (i.e. sinistral) is a much larger shell belonging to the Clausilia family. Thin, dark horn-colour, semi-transparent, glossy, 7 to 8 whorls, local, but abundant where found. Chiefly found on trees.
Clausilia (Pirostoma) bidentata.—All our British clausilias are sinistral. The clausilium (little door) is an internal contrivance fastened to the pillar of the shell (whereas an operculum is attached to the body of a mollusc) by an elastic ligament to protect it against insect enemies when the animal withdraws. Bidentata has two denticles, fusiform and reddish-brown, as are all. Very common on walls and trees.
Clausilia (Pirostoma) rolphii.—Rare and local. Almost subterranean in habit. More coarsely striated than the last. The upper whorls nearly of the same breadth, forming a short cylinder.
Clausilia (Alinda) biplicata.—Very local. Chiefly on Thames willows. Larger than the two former, and streaked with white.
Clausilia (Marpessa) laminata.—Much like the former, but widely distributed. Usually found on beech and ash trees, and on limestone rocks. Smooth and glossy.
In the family Stenogyra we have three genera, Azeca, Cochlicopa, and Cæcilioides (with also the imported Stenogyra Goodallii, found only in pine-houses).
Stenogyra (Azeca) tridens is a small chrysalis-shaped, solid but semi-transparent shell, horn-coloured, with 3 denticles. Not rare in moist places.
Stenogyra (Cochlicopa) lubrica (i.e. slippery).—Very common in moss and under stones or logs. Much like the previous species, but no denticles and fewer whorls, and broader mouth.
Stenogyra (Cæcilioides) acicula.—If this word is supposed to be Latin it would mean either “like to a blind worm” or “like to a lettuce”! Cæcus, however, being Latin for blind, the allusion is no doubt to the fact that this wholly subterranean species is eyeless. The only British representative of a large family of carnivorous molluscs. I have found it on Saxon bones when unearthed, and in crevices of limestone underground, but it is generally found dead amongst the rejectamenta on the banks of rivers. It is a pretty, glossy white shell, 5 millimetres in height by 1 in breadth.
I may notice here two other land shells, although they scientifically are grouped amongst the fluviatile Gasteropoda.
Cyclostoma (Pomatias) elegans.—Common on calcareous soils, especially chalk. A spiral shell of 4-1/2 whorls, suture very deep. Mouth circular (whence its name) and provided with a thick shelly operculum which closes the orifice when the animal retires by means of an elastic ligament. This and the next species are our only land shells provided with an operculum, and this shows their derivation from the marine Gasteropoda (e.g. whelk and winkle). Perhaps all shells were originally marine, but some became first amphibious and then terrestrial. It is quite unlike any other of our land shells.
Acicula lineata is a very small shell, the size of the Pupæ; mainly northern in distribution. Feeds on liverworts and fungi. Very local; 6 or 7 whorls. Mouth pear-shaped, with a horny operculum.
The Family Succinea really ranks with the land shells, as belonging to the sub-order Pulmonata or lung-breathing molluscs. It is, however, amphibious, and hibernates in the mud at the bottom of a ditch.
Succinea putris (it is the mud, not the animal, which is putrid!) is called the Amber Snail from the colour of its shell, which is unlike any other. Common on flags, etc., at the edges of ditches and ponds.
Succinea elegans.—Difficult to distinguish from the former, but the animal is darker and the shell more slender, with a deeper suture and a narrower mouth.
Succinea oblonga is local and rare. Generally found near the sea. Much smaller than the other Succineas, and easily mistaken for the young of other species. Colour dull greenish.
The family Auriculidæ is represented in Britain only by Carychium minimum; a very small, semi-transparent, white and glossy shell found under mossy stones and other moist places. Common, but sharp eyes are needed to find it.
We now come to the freshwater shells, which we capture best by means of a perforated scoop, whether they are on the waterweeds or hidden in the sand or mud of the bottom.
It may be noted that all freshwater shells are greenish-brown which is an excellent protective colouring as rendering them less visible among water weeds to the fish, which devour them greedily.
The family of Limnæidæ (or lake dwellers) has the sub-families, Planorbis, Physa, Limnæa, and Ancylus. In the Planorbinæ (i.e. flat-coiled) the only representative of the genus Segmentina is Segmentina nitida, a small, quoit-shaped, keeled, semi-transparent, light brown shell, with internal divisions like those of a nautilus which are visible from the outside of the shell. Local. Found in stagnant or sluggish water. The genus Planorbis contains the sub-genera Hippeutis, Gyraulus, Gyrorbis, Coretus, and Bathyomphalus.
Planorbis (Hippeutis) fontanus is much like Segmentina but has no septa, and is flatter. Common, especially on watercress. Often encrusted with mud.
Planorbis (Gyraulus) nautileus is very small; quoit-shaped, with the upper side flat. Grey and striated. The variety crista has the ridges of the epidermis drawn into points, and is beautiful when seen by a magnifying glass. Common in ponds and ditches.
Planorbis (Gyraulus) dilatatus is a very small shell imported in cotton bales from America, and naturalized in canals in Lancashire. No other of its kind is so small.
Planorbis (Gyraulus) albus is dull white and striated. Flattish above, with spire depressed. Frequently encrusted and black with mud. Common.
Planorbis (Gyraulus) parvus (but not so small as dilatatus).—Convex above with a central depression, concave beneath. Suture deep, and umbilicus large. Smooth and glossy. Local.
Planorbis (Gyrorbis) spirorbis.—Very flat, glossy, brown, whorls 5 to 6. Common in ponds and ditches.
Planorbis (Gyrorbis) vertex.—Very like the last, but flatter and thinner, and with a prominent keel. More local than spirorbis, but sometimes found with it. Whorls 6 to 8.
Planorbis (Gyrorbis) carinatus.—Larger than spirorbis and vertex. Sharply keeled in the centre of the outer margin. Mouth angulated above and below. Local, mainly in the south and east of England.
Planorbis (Gyrorbis) umbilicatus.—Like the last, but the keel is below and not on the centre. Mouth rhomboidal. More common than carinatus.
Planorbis (Coretus) corneus.—Far the largest species. Dark brown, lighter below. Mouth nearly circular. Spire sunk. In boiling water often exudes a crimson fluid. Common.
Planorbis (Bathyomphalus) contortus.—Small, 8-whorled, flat above, very convex below. Fairly common in still water. Very compact in appearance.
The sub-family Physa has two genera, Aplecta and Physa.
Physa (Aplecta) hypnorum is a spindle-shaped, very glossy, semi-transparent, dark reddish brown, shell, with 6 to 7 whorls. Not common. Found in still water.
Physa (Physa) fontinalis.—More common, and found in running as well as in still water. Shorter and more rounded than the last. Shell very thin, greenish horn-colour. Lobes of the mantle expand over the shell. Seen in an aquarium are its perpendicular threads of mucus, up and down which the animals climb.
Limnæa (Amphipeplea) glutinosa.—Very local. Somewhat like Ph. fontinalis, but larger and more thin. In young specimens the mantle covers the shell, and in adults the animal is not wholly contained in the shell.
Limnæa (sub-genus Radix) involuta.—Only found in one Irish tarn. Whorls envelop the spire. Very thin, pale amber.
Limnæa (Radix) peregra.—The most common and variable of all our freshwater shells. Spire pointed. Somewhat amphibious. Found practically over the whole of the Eastern Hemisphere.
Limnæa (Radix) auricularia.—Mouth very large, with outer lip widely reflected. Very common and fine in the Thames. Spire very short, apex sharp.
Limnæa (sub-genus Limnophysa) stagnalis.—The largest of the genus. Common, except in Wales. Shell greyish, spire long and tapering to a point; 12210 teeth on its lingual ribbon. See the illustration on p. 57, which also shows above two specimens of Paludina contecta, one being covered (as freshwater shells often are) by a vegetable growth, which obscures the marking.
Limnæa (Limnophysa) palustris.—Shell tapering, somewhat solid, brown, much smaller than stagnalis. Common in slow or stagnant water. Some varieties much darker than the type.
Limnæa (Limnophysa) truncatula.—Like the last in shape, but much smaller, and with a deeper suture. Common, and fond of being out of the water. A parasite of this mollusc causes “fluke” in sheep which have taken it in by drinking or by eating grass by the side of ponds and ditches.
Paludina contecta (two) and Limnæa stagnalis on water-weeds.
Limnæa (Omphiscola) glabra.—Also amphibious. About the same size as truncatula. Local. Inner lip rather thick and reflected on the base of the penultimate whorl.
Limnæa (Ancylus) fluviatilis.—“Freshwater limpet.” Shell, rather limpet-like, with a hooked apex (whence its generic name), adheres to stones or piles in running water. Common. I once dredged a large water-beetle with three of these shells adhering to its wing-cases; thus it would be transported to fresh habitats.
Limnæa (Acroloxus) lacustris.—Like the former but more local, and preferring sluggish or still waters. Shell more oblong, thinner, and apex twisted to the left instead of to the right as in fluviatilis.
The sub-order Pectinibranchiata (comb-like gill) contains the genera Neritina, Paludina, and Valvata, in all of which there are two tentacles with eye at the base, and an operculum to the shell.
Neritina fluviatilis.—Solid, glossy, chequered brown, white, and purple (but also a lemon-coloured variety). Operculum semi-lunar, orange, with a projection which serves as a lock to keep the operculum in position. Not rare in England; on stones in running water. See illustration below, which also shows above L. (Ancylus) fluviatilis.
Paludina (Vivipara) contecta.—Shell dark green with darker bands. Conical. Suture very deep. Operculum horny. Viviparous. Local.
Paludina (Vivipara) vivipara.—More common than contecta. Shell more oval, not so glossy, light greenish yellow, suture not so deep, no umbilicus, apex blunt.
Neritina and Ancylus.
Paludina (Bythinia) tentaculata.—(The eyes in this genus are not on foot-stalks; the operculum is shelly instead of horny). Common in slow water and ditches. Shell semi-transparent, yellowish, mouth oval, angulated above. Operculum made of plates rising one above another formed at different stages of growth.
Paludina (Bythinia) leachii.—Much smaller and less common than the last. Distinct umbilicus; mouth almost circular.
Paludina (Paludestrina) ventrosa.—A brackish-water shell, swarming where found, e.g. from Erith to Gravesend, and in East Anglia. Shell small, thin, semi-transparent.
Paludina (Paludestrina) jenkinsi.—A larger shell, not confined to brackish water and spreading very rapidly. Swarms where found. A variety has a marked keel which sometimes bears bunches of spines at equal distances.
Paludina (Paludestrina) stagnalis.—Larger and with more whorls. Not so common.
Paludina (Pseudamnicola) anatina.—Small, sub-conical, deep suture. Found in brackish water, and apparently identical with Hydrobia or Paludestrina similis, which I used to find by the Thames, where it is now apparently extinct.
Valvata piscinalis.—Globular, suture very deep, circular mouth, operculum concentrically spiral. In ponds and slow water. Shell yellowish, but commonly covered with conferva.
Valvata cristata.—Much smaller; shell disk-shaped. Frequents the roots of flags. Shell striated and more or less ridged, but the name cristata refers to the plume-like appearance of its breathing apparatus.
We now come to the bivalve shells with leaf-like gills. The Unionidæ contain two genera, Unio and Anodonta, commonly called freshwater mussels.
Unio tumidus.—Shell ovate, very solid, dark brown; common. See accompanying illustration, which shows the fringed branchial siphon which draws in food-bearing water, and the smaller anal siphon by which it gets rid of undigested matter.
Freshwater mussel breathing and eating.
Unio pictorum.—More oblong and thinner shell, yellowish, girdled with brown in the lines of growth. Common. The specific name recalls that gold and silver paint used to be sold in these shells (or marine mussels) for illuminating work. It is said to produce 220,000 eggs in the three summer months.
Unio (margaritana) margaritifer.—Shell solid and black, beaks always eroded. Mainly found in mountain streams. Its pearls are few and poor compared with those of marine shells; but they attracted the notice of Cæsar and so hastened the conquest (and development) of Britain.
Anodonta cygnea.—(In this genus the hinge is toothless, whence its generic name. The specific names cygnea and anatina mean “swan” and “duck,” in reference to their comparative size). This is the largest of our freshwater shells, reaching even 9 inches in breadth by 4-1/2 in length. Common in ponds and slow water. Sometimes the shells are yellowish green with rays of the same colour.
Anodonta anatina.—Doubtful if this is a separate species or only a smaller form. The hinge line is raised instead of being straight, and the posterior side slopes abruptly instead of gradually.
In the next family are two genera, Sphærium and Pisidium.
Sphærium corneum.—Very common. Shell somewhat globular, glossy, opaque, horn-coloured, marked with lighter bands in the line of growth. Usually on the bottom, but can suspend itself by threads of mucus.
Sphærium rivicola.—Much larger. Also flatter and more striated. Yellowish brown or greenish. A whole series of young of different sizes will be found in the animal.
Sphærium pallidum.—Local in canals and ponds. Oblong. Distinguished also from the previous species by the body being milk-white, and the shell is ashy-grey.
Sphærium lacustre.—Local. On the beaks is a calcareous nucleus which distinguishes it. It is thinner than corneum, and rounder than pallidum.
Pisidium amnicum.—(Our five pisidia resemble Sphærium, but are much smaller, all but amnicus being minute. Very abundant where found. P. amnicum and fortinale are triangular in shape, P. pusillum oval, P. nitidum round, and P. roseum or milium oblong; but they are difficult to distinguish on account of their similarity and variation). P. amnicum is nearly twice the size of the others, and this and fontinale may be found in slow rivers, whereas the others prefer stagnant waters.
Pisidium fontinale.—Smaller and thinner, and with more prominent beaks than P. amnicum.
Pisidium pusillum.—The most common species. Distinguished from the last by being oval and by its beaks being blunter and more central.
Pisidium nitidum.—Rare. Very glossy and striated.
Pisidium roseum (from the colour of part of its body).—Like nitidum, but oblong, with a straight lower margin, and with beaks placed away from the centre.
The last shell to be mentioned could not be mistaken for any other. It belongs to the sub-order Heteromya (i.e. with adductor or closing muscles not equal); to the family of Mytilidæ (or mussels) and the genus Dreissensia (named after a Dutch conchologist).
Dreissensia polymorpha is a triangular, boat-shaped, bivalve, supposed to have been introduced with Russian timber (as was also probably Hydrobia Jenkinsi). It is gregarious, and attaches itself to objects by a byssus like our marine mussels. Shell yellowish-brown with wavy purplish lines, wrinkled in the line of growth. Common in the New River, and has been found in iron water-pipes in Oxford Street.
All our shells have varieties (many an albino or white form), and the collection and distinguishing of these varieties, which in some species are numerous, adds much to the interest of the collector. In addition there are also the variations in size or markings which can hardly rank as varieties. Inasmuch as none of our shells are peculiar to our country (which is from the natural history and the geological point of view only a detached portion of the Continent), it may be well to warn young collectors that if they receive shells from the Continent, mere varieties are there often named as separate species and variations considered as definite varieties. This is especially the case with Helicella virgata.
As to the arrangement of shells in a collection before a regular cabinet is obtained, the tinier shells may be kept in small glass tubes with corks (such as used for homœopathic medicines), and the medium sized ones in the trays of common matchboxes, these being arranged in large shallow glass-covered trays which can be obtained from any cardboard boxmaker at a small cost, and several of these, stored one above the other, form an excellent substitute for a more costly cabinet. In all cases the name, and the place where the shells were found, should be written on a small slip of card placed in the tube or tray. It is not well in most cases to fasten the shells on card, but if this is done gum tragacanth is best. The collection should be kept free from damp and from dust.
The following notes supply a few general rules as to finding and preserving shells:—
Of Shell-bearing Molluscs there are three classes—Marine, Freshwater, and Land. The first two include Univalves and Bivalves, the last only Univalves.
1. Marine Shells may be obtained, 1st, by searching on and under rocks at low water, or on coral reefs, among seaweed attached to them, or floating on the sea, or on a sandy beach. Bivalves may be found by digging in the sand, or mud, on a beach, or at the mouth of a river: their presence is generally indicated by a circular breathing hole in the sand. 2nd. By dredging, by which means only deep-sea shells can be obtained; but after a storm these may often be found upon the shore, before they have lost their lustre.
Limpets, etc., should be detached with a thin blade passed quickly under the shell, taking care not to break the edges. Small shells on and in seaweed, and limpets, etc., adhering to stones will drop off and sink to the bottom in a vessel of cold fresh water.
2. Fresh-water Shells may be obtained in any river, lake, pond, marsh or reservoir. Univalves, chiefly on the banks, on reeds and plants growing near the hedges, and on the under surface, leaves, and stems of aquatic plants. Bivalves generally at the bottom, among stones, or buried in the sand, or among the roots of aquatic plants.
3. Land Shells.—These resemble, more or less, in their habits the garden snail, though varying greatly in character, size, and colour. They mostly abound in a chalk or limestone district, and in moist and wooded situations. Some species inhabit low and damp spots, roots of trees, hollows and crevices of rocks and walls; some lie under stones or pieces of wood, or in the earth; others climb shrubs, and in tropical climates even lofty trees. Their haunts vary according to the weather and the season. They come out early in the morning, and after rain. Some bury themselves in moist places during the dry season, or burrow under leaves, grass, or stones, often closing the mouths of their shells with a white secretion to prevent evaporation during the period of hibernation.
The smallest shells, especially of land species, and young imperfect shells should be collected.
In all cases “live shells,” i.e. shells in which the animal is alive, are to be chosen; but, when these cannot be procured, “dead shells,” which have not lost their lustre, or their colour, especially those of rare species, should be preserved.
1. No attempt should be made to clean them, or to remove the furry skin, more or less thick, with which they are often covered, beyond removing with a soft brush any mud or sand adhering to them.
2. The animals of Land and Freshwater shells may be killed by immersing them for a few minutes in boiling water, after which the bodies may be easily extracted whole with any suitable instrument, e.g., a fork or a pin, according to size. Hot water should not be used with marine shells: it often destroys their lustre. They should be buried, if time permits, in sand, or other dry material, until the animal dries up (in small shells) or rots (in large specimens); or they may be drowned in cold fresh water, and hung up in the air to dry or rot away. In the former case, if an operculum (with which some species, both marine and land, close their mouths, more or less partially) exists, it will, generally in the case of land shells, remain in its place, adhering to the shell. In the latter, the decayed matter should be washed out, and the operculum, if any, replaced and fixed, say, on cotton filling the shell. This applies equally to land shells.
3. Care should be taken not to injure the edge or lip of the mouth of univalves, or the ligament of the hinge of bivalves. When bivalves gape on dying in water, or if the ligament be broken, the valves should be closed and tied together. If the ligament of a gaping bivalve should become dry and stiff, it can be softened by putting it in water.
4. The localities in which each species is found should be noted, and, in the case of dredging, the depth of water.
All solid shells may be wrapped in one or two folds of paper of any kind. Fragile and minute shells should be put, generally separately, into a box or bottle—with or without cotton, as required. Such packets may be heaped up in any box, heavy shells at the bottom, without pressure, and any blank filled at the top with paper or other elastic material. Sawdust injures the lustre of many species.
Two books on shells should be procured at an early stage of the collector’s career, which will give not only minute descriptions of all our land and freshwater shells and their varieties, but also plates of illustrations. These are the Collector’s Manual, by L. E. Adams, 2nd ed., published by Taylor Brothers of Leeds; and Rimmers’ Land and Fresh Water Shells, published by George Grant of Edinburgh.
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