From owner-biophysics@net.bio.net Tue Mar 30 23:00:00 1999 From: "Clive Delmonte" Newsgroups: bionet.biophysics,bionet.xtallography Subject: DNA Structure: Puzzle #7 (Version 2) Date: Wed, 31 Mar 1999 17:08:12 +0100 Lines: 1031 X-Newsreader: Microsoft Outlook Express 4.72.3110.5 X-MimeOLE: Produced By Microsoft MimeOLE V4.72.3110.3 NNTP-Posting-Host: 195.7.225.41 Message-ID: <37024616.0@news.netdirect.net.uk> X-Trace: 31 Mar 1999 16:58:14 GMT, 195.7.225.41 Path: biosci!news.ic.sunysb.edu!news-nysernet-16.sprintlink.net!news-east1.sprintlink.net!news-peer1.sprintlink.net!news.sprintlink.net!news-peer.gip.net!news-lond.gip.net!news.gsl.net!gip.net!newspeer.clara.net!news.clara.net!peer.news.nildram.co.uk!news-peer.netdirect.net.uk!news.netdirect.net.uk!195.7.225.41 Xref: biosci bionet.biophysics:4898 bionet.xtallography:4682 The Unwinding Problem Inside Fibres Leslie et al.(19) reported that a fibre of poly(dI).poly(dC), in the absence of unwinding proteins, which gave sharp X-ray diffraction spots shortly after being drawn, was unstable and transformed irreversibly after a few days into poly(dC).poly(dI).poly(dC+). This new, three-stranded molecule also gave sharp spots in its own X-ray diffraction pattern which did not have any of the original spots. Therefore the conversion was complete. Very similar results had been reported for fibres of poly(dA).poly(dT) converting to poly(dT).poly(dA).poly(dT) (20) and for the formation of poly(U).poly(A).poly(U) from poly(U).poly(A) (21). The problem is evident: How and where would the torque arise, and be sustained, inside a solid fibre that would permit one double helix to rotate so as to unwind one strand and rewind it onto an adjacent triple helix having a different diameter and rotating at a different angular velocity, the conversion being complete ? Inside the fibre, individual molecules would have a random axial translation so only very rarely would two adjacent double helices have the same axial starting point down the fibre, and they would be unlikely to be perfectly straight inside the fibre but would probably be entwined around neighbours. A true side-by-side duplex DNA structure (see attachment) could form a side-by-side triplex, without any unwinding, by the linear transfer of a strand across to a neighbouring duplex. ----------------------------------------------------------------- 19 Polymorphism of DNA Double Helices; A.G.W. Leslie et al.; J Mol Biol Vol 143 (1980) 49 - 72 20 Structures for the Polynucleotide Complexes Poly(dA).Poly(dT) & Poly(dT).Poly(dA).Poly(dT); Struther Arnott & E. Selsing; J Mol Biol Vol 88 (1974) 509-521 21 Structures for Poly(U).poly(A).poly(U) Triple Stranded Polynucleotides; Struther Arnott & P.J. 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!____^```!___\?___\?___[____W_____ M_______\_P__\P?_________________\ #______________]__?____^?_ M____\_____S___^____#_____________ `/__/ __________________ ` M_______________?_W_____X_____Q______?___@ #_'_____________\` M___SX__________________P`/_______________________P?___!_____ M_X___[___'__________________<^____________________Q__/______________ M___P`/_________\__?_X?#________'/C^.+^A\./XS['P_\G______ M______________^ `__________________P`/_________\!___WW]_____ M___'CB_H?#C^,^Q\/_)_____________________@ /_________________ M\ #__________(#__]]_?_______QXX_\'PX_C_P?#YS________________ M______ ?__________________ `___________8'__??W_______\>./O/\ M./X^\/\8<^_________________________________________P`/______ M____WP?_P !_______\`/ #[^ '\`/S_QD /________________________ M________________\ #__________]_/_\ `?_______________________ M_____________________P_@__________________ `___________?S__ M`'____________________________________________\/X/__________ M_______P`/__________W\__P !_________________________________ M____________#^#_________________\ #__________-@__]__?_______ M_____________________________________W_\__________________ ` M__________R'___?_W__________________________________________ M__]P'/_________________P`/_________\?_______________________ M____________________________?WS_________________\ #_________ M_'___________________________________________________W]\____ M______________ `__________S_________________________________ M__________________]_?/_________________P`/__________________ M____________________________________________``#_____________ M____\ #_____________________________________________________ M_________P``__________________ `____________________________ M__________________________________]__/_________________P`/__ M____________________________________________________________ M?_S_________________\ #_____________________________________ M_________________________W_\__________________ `____________ M__________________________________________________]__/______ M___________P`/______________________________________________ M____________________________________\ #_____________________ M____________________________________________________________ M__ `________________________________________________________ M___________________________P`/______________________________ M____________________________________________________\ #_____ M____________________________________________________________ M__________________ `________________________________________ M___________________\?______________________P`/______________ M____________________________________________________________ M________\ #_________________________________________________ M__________________________________ `________________________ M___________________________________________________________P M`/__________________________________________________________ M________________________\ #_________________________________ M__________________________________________________ `________ M____________________________________________________________ M_______________P`/__________________________________________ M________________________________________\ #_________________ M____________________________________________________________ M______ `____________________________________________________ M_______________________________P`/__________________________ M________________________________________________________\ #_ M____________________________________________________________ M______________________ `____________________________________ M_______________________________________________P`/__________ M____________________________________________________________ M____________\ #_____________________________________________ M______________________________________ `____________________ M____________________________________________________________ M___P`/______________________________________________________ M____________________________\ #_____________________________ M______________________________________________________ `____ M____________________________________________________________ M___________________P`/______________________________________ M____________________________________________\ #_____________ M____________________________________________________________ M__________ `________________________________________________ M___________________________________P`/______________________ M____________________________________________________________ M\ #_________________________________________________________ M__________________________ `________________________________ M___________________________________________________P`/______ M____________________________________________________________ M________________\ #_________________________________________ M__________________________________________ `________________ M____________________________________________________________ M_______P`/__________________________________________________ M________________________________\ #_________________________ M__________________________________________________________ ` M____________________________________________________________ M_______________________P`/__________________________________ M________________________________________________\ #_________ M____________________________________________________________ M______________ `____________________________________________ M_______________________________________P`/__________________ M____________________________________________________________ M____\ #_____________________________________________________ M______________________________ `____________________________ M_______________________________________________________P`/__ M____________________________________________________________ M____________________\ #_____________________________________ M______________________________________________ `____________ M____________________________________________________________ M___________P`/______________________________________________ M____________________________________\ #_____________________ M____________________________________________________________ M__ `________________________________________________________ M___________________________P`/______________________________ M____________________________________________________\ #_____ M____________________________________________________________ M__________________ `________________________________________ M___________________________________________P`/______________ M____________________________________________________________ M________\ #_________________________________________________ M__________________________________ `________________________ M___________________________________________________________P M`/__________________________________________________________ M________________________\ #_________________________________ M__________________________________________________ `________ M____________________________________________________________ M_______________P`/__________________________________________ M________________________________________\ #_________________ M____________________________________________________________ M______ `____________________________________________________ M_______________________________P`/__________________________ M________________________________________________________\ #_ M____________________________________________________________ M______________________ `____________________________________ M_______________________________________________P`/__________ M____________________________________________________________ M____________\ #_____________________________________________ M______________________________________ `____________________ M____________________________________________________________ M___P`/______________________________________________________ M____________________________\ #_____________________________ M______________________________________________________ `____ M____________________________________________________________ M___________________P`/______________________________________ M____________________________________________\ #_____________ M____________________________________________________________ M__________ `________________________________________________ M___________________________________P`/______________________ M____________________________________________________________ M\ #_________________________________________________________ M__________________________ `________________________________ M___________________________________________________P`/______ M____________________________________________________________ M________________\ #_________________________________________ M__________________________________________ `________________ M____________________________________________________________ M_______P`/__________________________________________________ M________________________________\ #_________________________ M__________________________________________________________ ` M____________________________________________________________ M_______________________P`/__________________________________ M________________________________________________\ #_________ M____________________________________________________________ M______________ `____________________________________________ M_______________________________________P`/__________________ M____________________________________________________________ M____\ #_____________________________________________________ M______________________________ `____________________________ M_______________________________________________________P`/__ M____________________________________________________________ M____________________\ #_____________________________________ M______________________________________________ `____________ M____________________________________________________________ M___________P`/______________________________________________ M____________________________________\ #_____________________ M____________________________________________________________ M__ `________________________________________________________ M___________________________P`/______________________________ M____________________________________________________\ #_____ M____________________________________________________________ M__________________ `________________________________________ M___________________________________________P`/______________ M____________________________________________________________ M________\ #_________________________________________________ M__________________________________ `________________________ M___________________________________________________________P M`/__________________________________________________________ M________________________\ #_________________________________ M__________________________________________________ `________ M____________________________________________________________ M_______________P`/__________________________________________ M________________________________________\ #_________________ M____________________________________________________________ M______ `____________________________________________________ M_______________________________P`/__________________________ M________________________________________________________\ #_ M____________________________________________________________ M______________________ `____________________________________ M_______________________________________________P`/__________ M____________________________________________________________ M____________\ #_____________________________________________ M______________________________________ `____________________ M____________________________________________________________ M___P`/______________________________________________________ M____________________________\ #_____________________________ M______________________________________________________ `____ M____________________________________________________________ M___________________P`/______________________________________ M____________________________________________\ #_____________ M____________________________________________________________ M__________ `________________________________________________ M___________________________________P`/______________________ M____________________________________________________________ M\ #_________________________________________________________ M__________________________ `________________________________ M___________________________________________________P`/______ M____________________________________________________________ M________________\ #_________________________________________ M__________________________________________ `________________ M____________________________________________________________ M_______P`/__________________________________________________ M________________________________\ #_________________________ M__________________________________________________________ ` M____________________________________________________________ M_______________________P`/__________________________________ M________________________________________________\ #_________ M____________________________________________________________ M______________ `____________________________________________ M_______________________________________P`/__________________ M____________________________________________________________ M____\ #_____________________________________________________ M______________________________ `____________________________ M_______________________________________________________P`/__ M____________________________________________________________ M____________________\ #_____________________________________ M______________________________________________ `____________ M____________________________________________________________ M___________P`/______________________________________________ M____________________________________\ #_____________________ M____________________________________________________________ M__ `________________________________________________________ M___________________________P`/______________________________ M____________________________________________________\ #_____ M____________________________________________________________ M__________________ `________________________________________ M___________________________________________P`/______________ M____________________________________________________________ M________\ #_________________________________________________ ;__________________________________ ` ` end .