From owner-biophysics@net.bio.net Tue Mar 09 22:00:00 1999 From: "Clive Delmonte" Newsgroups: bionet.biophysics,bionet.xtallography Subject: DNA Structure: Puzzle #2 (Version 2) Date: Wed, 10 Mar 1999 20:14:56 -0000 Lines: 1976 X-Newsreader: Microsoft Outlook Express 4.72.3110.5 X-MimeOLE: Produced By Microsoft MimeOLE V4.72.3110.3 NNTP-Posting-Host: 195.7.225.115 Message-ID: <36e6cf64.0@news.netdirect.net.uk> X-Trace: 10 Mar 1999 20:00:36 GMT, 195.7.225.115 Path: biosci!news.ic.sunysb.edu!news-nysernet-16.sprintlink.net!news-east1.sprintlink.net!news-peer1.sprintlink.net!news.sprintlink.net!news-peer.gip.net!news-lond.gip.net!news.gsl.net!gip.net!newspeer.clara.net!news.clara.net!peer.news.nildram.co.uk!news-peer.netdirect.net.uk!news.netdirect.net.uk!195.7.225.115 Xref: biosci bionet.biophysics:4853 bionet.xtallography:4646 Optical Diffraction Compared With X-Ray Diffraction for B-DNA It was Franklin's famous diffraction photograph of B-DNA (3), shown to him by Wilkins, that Watson sketched on the train back to Cambridge to rejoin Crick. One of Wilkins' collaborators, A R Stokes (4), produced probably the first optical diffraction prediction of the pattern arising from a true side-by-side helical structure, this one having 8 repeat units per full turn, a montage of which I have assembled as attachment 1. This true side-by-side structure mirrors that portrayed in attachment 2 (taken from ref. 2 (Puzzle #1)). The likeness of Stokes' optical diffraction pattern would seem to resemble closely Franklin's X-ray diffraction pattern (attachment 3). In the same paper (4), Stokes, from his work on the Fourier transform for a helix, showed that, defining as "delta" the angle between an arm of the "helical cross" and the Y-axis (meridian), tan delta = helical pitch / helical circumference Applying this result to Franklin's photograph of B-DNA (see attachment 3), delta is measured as 40 degrees, and, with a pitch of 3.34 nm, the helical diameter is found to be some 1.3 nm. Even in later work with better definition, for example, from Wilkins' group (attachment 4), we see a similar value of delta. For a supposed double helical diameter of 2 to 2.2 nm, delta should be 28 - 26 degrees respectively, far removed from the actual values of 40 - 41. The helical diameter of 1.3 nm, found from Stokes' equation applied to Franklin's & Wilkins' photographs, coincides with the STM value of 1.3 nm measured by Lee et al. (Puzzle #1, ref. 1), recorded in their Figure 3(D), and accords with the Watson-Crick base pair width of 1.1 nm which determines the sugar-phosphate chain diameter set out in the attachment 2 here. ------------------------------------------------------------- 3 Molecular Configuration in Sodium Thymonucleate; R.E. Franklin & R.G. Gosling; Nature Vol 171 (1953) 740 - 741 4 The Theory of X-ray Fibre Diagrams; A.R. Stokes; Prog. Biophys. & Biophys. Chem. 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